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Douglas' bluegrass, sand dune blue grass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; densely to loosely tufted or with solitary shoots, sometimes stoloniferous, sometimes rhizomatous.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

(5)10-125 cm, terete or weakly compressed;

nodes terete or slightly compressed.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed from 1/7 their length to their entire length, terete to compressed, smooth or scabrous, glabrous or pubescent;

ligules 0.2-18 mm, milky white or colorless, usually translucent, truncate to acuminate, glabrous or ciliolate;

innovation blades with the adaxial surfaces usually moderately to densely scabrous or hispidulous on and between the veins, sometimes smooth and glabrous;

cauline blades flat, folded, or involute, thin or thick, lax or straight, smooth or scabrous, adaxial surfaces sometimes hairy, apices narrowly to broadly prow-shaped.

Basal branching

mainly intravaginal, some extravaginal.

intra- and/or extravaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

1-29 cm, contracted to open, usually with fewer than 100 spikelets;

nodes with 1-5 branches;

branches 0.5-18 cm, terete or angled, smooth or scabrous, glabrous or hispidulous.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

3-17 mm, lengths 3.5 times widths, laterally compressed, not sexually dimorphic, not bulbiferous;

florets 2-10(13) mm, normal;

rachilla internodes smooth or scabrous, glabrous or hairy.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

distinctly keeled, keels smooth or scabrous;

lower glumes 1, 3(or 5)-veined;

upper glumes 3- or 5-veined;

calluses terete or slightly laterally compressed, glabrous, webbed, or with a crown of hairs;

lemmas 2.6-11 mm, lanceolate, distinctly keeled, keels, veins, and intercostal regions glabrous or hairy, 5-7(11)-veined;

palea keels scabrous, glabrous or with hairs at midlength;

anthers 3, vestigial (0.1-0.2 mm) or 1.3-4.5(5) mm.

2n

= 28.

Poa douglasii

Poa sect. Madropoa

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa sect. Madropoa is confined to North America. Its 20 species exhibit breeding systems ranging from sequential gynomonoecy to gynodioecy and dioecy. The gynomonoecious species usually grow in forests and have broad, flat leaves. The gynomonoecious and dioecious species grow mainly in more open habitats. They have normally developed anthers that are 1.3-4 mm long, and involute innovation blades that, in several species, are densely scabrous or hairy on the adaxial surfaces.

There are two subsections in the Flora region: subsects. Madropoa and Epiles.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 551. FNA vol. 24.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
Name authority Nees Soreng
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