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Douglas' bluegrass, sand dune blue grass

large-glume bluegrass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; green or strongly purplish; rhizomatous, rhizomes sometimes poorly developed, densely to loosely tufted.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

(20)30-120 cm, usually stout, erect or decumbent, not branching above the base, terete or weakly compressed;

nodes terete, 1-2 exserted.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed for 1/4 - 3/5 their length, terete to compressed, usually distinctly keeled, glabrous, sparsely to moderately scabrous, bases of basal sheaths glabrous, distal sheath lengths (0.8)1-2(2.5) times blade lengths;

collars smooth, glabrous;

ligules 2-5(6) mm, usually smooth, ligules of proximal leaves sometimes scabrous, obtuse, those of the distal leaves acute;

blades 4-12(18) cm long, (2)3-7 mm wide, flat or loosely folded, glabrous, both surfaces smooth or the adaxial surfaces sparsely scabrous, apices broadly prow-shaped, cauline blades subequal, flag leaf blades 3-10 cm.

Basal branching

mainly intravaginal, some extravaginal.

all or mainly extravaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

(4)7-15(20) cm, lax, loosely contracted or open, moderately congested, with (15)20-100 spikelets and 2-4(5) branches per node;

branches 3-8 cm, ascending or eventually spreading, flexuous, terete or weakly angled, usually sparsely scabridulous or scabrous, infrequently moderately densely scabrous, with (3)5-15(30) spikelets moderately crowded in the distal 1/2.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

(5)6-9(12) mm, lengths to 3 times widths, laterally compressed, sometimes bulbiferous;

florets 2-5, infrequently bulb-forming;

rachilla internodes smooth or sparsely hispidulous.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

subequal in length and width, usually nearly equaling the adjacent lemmas, lanceolate to broadly lanceolate, distinctly keeled, keels sparsely to moderately scabrous distally, lateral veins prominent;

lower glumes (4)4.5-7 mm, 3-veined: upper glumes (4)4.5-7(8) mm, 3(5)-veined;

calluses dorsally webbed, hairs 1/3 – 2/3 times the lemma length, copious;

lemmas (4)5-8 mm, lanceolate to broadly lanceolate, green or purple, distinctly keeled, keels and marginal veins long-villous, lateral veins prominent, usually softly puberulent to short-villous, intercostal regions sometimes hispidulous, infrequently softly puberulent or short-villous proximally and hispidulous in the central 1/3, sometimes merely muriculate proximally and scabrous distally, sometimes densely scabrous throughout, rarely glabrous and nearly smooth proximally and sparsely scabrous distally, margins broadly hyaline, glabrous, apices acute;

palea keels scabrous, sometimes softly puberulent at midlength, intercostal regions broad, distinctly hispidulous or softly puberulent;

anthers 1.5-2.5 mm.

2n

= 28.

= 42, 43, 44, 45, ca. 46, 49, 56, ca. 58, 63, ca. 64, ca. 65, 66, 67, 68, 69, 70, ca. 71, ca. 72, ca. 73, 74, 75, 76, 77, 78, 80, ca. 82, ca. 84, ca. 87, ca. 100.

Poa douglasii

Poa macrocalyx

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; BC
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa macrocalyx grows mainly in coastal areas of boreal Alaska, and from the eastern coast of Russia to northern Japan. Bulbiferous plants are occasionally found. Poa macrocalyx resembles an exaggeratedly robust P. pratensis (p. 522), with large spikelets and lemmas, proportionally longer glumes, and paleas that are generally hispidulous between the veins and palea keels. It is cytologically and morphologically complex, and is sometimes difficult to distinguish from P. arctica subsp. lanata (see below). Poa norbergii Hulten may belong to this species; alternatively, it may be a hybrid between P. glauca (p. 576) and P. arctica (p. 576) or P. macrocalyx.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 551. FNA vol. 24, p. 527.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Poa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. turneri, P. ×norbergii, P. hispidula
Name authority Nees Trautv. & CA. Mey.
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