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Douglas' bluegrass, sand dune blue grass

dune bluegrass, large flower sand dune blue grass, sand-dune blue grass, seashore blue grass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 4 m, stout, robust.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

(7)15-60 cm tall, 1.5-2 mm thick, bases decumbent, terete or weakly compressed, smooth or moderately scabrous below the panicles;

nodes terete, 0(1) exserted.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed for about 1/2 their length, terete, glabrous or sparsely retrorsely scabrous, bases of basal sheaths glabrous, distal sheath lengths 1.7-4(6) times blade lengths;

collars smooth, glabrous;

ligules 1-5 mm, scabrous, truncate to acute, ciliolate;

innovation blades to 30 cm, moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 2-4 mm wide, involute, thick, somewhat arcuate, firm, abaxial surfaces smooth or moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-10 cm.

Basal branching

mainly intravaginal, some extravaginal.

mostly intravaginal, some extravaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

3-15 cm, erect, ovoid to lanceolate, contracted, often interrupted, congested, with 15-80 spikelets, rachises glabrous, smooth to moderately scabrous;

nodes with 1-2 branches;

branches 1-6 cm, erect, stiff, terete to weakly angled, smooth or sparsely to moderately scabrous, with 3-17 spikelets.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

9-17 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6(10);

rachilla internodes smooth, usually hairy, hairs 0.3-0.4+ mm, rarely glabrous.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

broadly lanceolate, subequal to the adjacent lemmas, distinctly keeled, keels sparsely scabrous near the apices;

lower glumes 3-veined;

upper glumes usually 7+ mm, 3-5-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas (6)7.5-11 mm, lanceolate, 5-7(11)-veined, distinctly keeled, keels and marginal veins, and sometimes the lateral veins, short-villous to softly puberulent, intercostal regions smooth or scabrous, glabrous or softly puberulent, margins glabrous, apices acute;

palea keels scabrous, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)3-4(5) mm.

2n

= 28.

= 28.

Poa douglasii

Poa macrantha

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa macrantha is a dioecious coastal sand dune species that grows from southern Alaska to northern California. It competes better than P. douglasii (see previous) with the invasion of its habitat by Ammophila and other exotic species. It used to be treated as a subspecies of P. douglasii; a few intermediates with that species have been found around the mouth of Little River, California. Although clearly related, the two species are reasonably divergent in a number of characters. Poa macrantha is readily distinguished from P. douglasii by its glabrous rachises and usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 551. FNA vol. 24, p. 551.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. douglasii subsp. macrantha
Name authority Nees Vasey
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