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Douglas' bluegrass, sand dune blue grass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; not or only slightly glaucous; densely tufted, not stoloniferous, not rhizomatous.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

6-18 cm, with 0(1) exserted nodes, upper node in the lower 1/3 of the culms.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed for 1/10 – 1/5 their length, terete, smooth or very sparsely scabrous, distal sheath lengths 0.8-1.8 times blade lengths;

ligules 1.25-2.5 mm, smooth, apices obtuse, often lacerate;

blades thin, sparsely scabrous adaxially, flag leaf blades 1.6-3.8 cm.

Basal branching

mainly intravaginal, some extravaginal.

mixed intra- and extra vaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

1.5-3.5 cm, slightly lax, ovoid, contracted to loosely contracted, dense to moderately dense, with 2-6 branches per node;

branches steeply ascending, fairly straight, sulcate or angled, smooth or infrequently the angles sparsely scabrous, not glaucous.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

laterally compressed;

florets 2-5;

rachilla internodes smooth, glabrous, lower internodes 0.8-1 mm.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

equal, broadly lanceolate, thin;

lower glumes 0.75-1.05 mm wide, 3-veined;

upper glumes 3.7-4.7 mm long, 0.9-1.3 mm wide, lengths 3.7-4.1 times widths;

calluses all glabrous, or some proximal florets within a spikelet sparsely webbed;

lemmas 3.7-4.5 mm, broadly lanceolate, distinctly keeled, thin, keels and marginal veins short- to long-villous, hairs extending 1/3-1/2 the keel length, lateral veins usually glabrous, or infrequently sparsely softly puberulent, intercostal regions glabrous;

palea keels finely scabrous;

anthers 0.8-1.2 mm, poorly formed, sacs not fully maturing, not dehiscing, about 0.1 mm in diameter.

2n

= 28.

= ca. 65, 70.

Poa douglasii

Poa laxa × glauca

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa laxa × glauca is an eastern low arctic entity which has passed under the name P. flexuosa Sm., P. laxa subsp. flexuosa (Sm.) Hyl., and, more recently, P. laxiuscula (Blytt) Lange. It has also been confused with P. glauca (p. 576). It can be distinguished from P. laxa (see previous) by its more open sheaths and poorly developed, indehiscent anthers. It differs from P. glauca in its broad, thin glumes and lemmas; compact panicles; smooth or nearly smooth, non-glaucous branches; and poorly developed, indehiscent anthers. It also grows in wetter habitats than P. glauca, often around seeps. Its chloroplast DNA is more like that of the American P. laxa subsp. fernaldiana than that of the European subspp. flexuosa and laxa or of P. glauca.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 551. FNA vol. 24, p. 572.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Oreinos
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Name authority Nees unknown
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