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Douglas' bluegrass, sand dune blue grass

banff bluegrass, glaucous bluegrass, lax bluegrass, Mt. Washington bluegrass, wavy blue grass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; not or only slightly glaucous; densely tufted, not stoloniferous, not rhizomatous.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

8-35 cm tall, 0.5-0.9 mm thick, ascending to erect, slender;

nodes terete, 0(1) exserted.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed for 1/5-1/3 their length, terete, smooth, glabrous, bases of basal sheaths glabrous;

collars smooth or scabrous, glabrous;

ligules 2-4 mm, smooth, apices acute, often lacerate;

innovation blades similar to the cauline blades;

cauline blades 1-2(3) mm wide, flat, thin, soft, smooth, narrowly prow-tipped, blades not strongly graduated or reduced upwards.

Basal branching

mainly intravaginal, some extravaginal.

mixed, mainly extravaginal or mainly pseudointravaginal, sometimes intravaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

2-8 cm, slightly lax, usually loosely contracted and sparse, infrequently contracted and dense;

nodes with 1-3(5) branches;

branches 1-3(4) cm, usually ascending or weakly spreading, infrequently erect, fairly straight or flexuous, slender, sulcate or angled, smooth or the angles sparsely scabrous, with 1-8 spikelets.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

4-6 mm, lengths to 3 times widths, laterally compressed;

florets 2-5;

rachilla internodes shorter than 1 mm, smooth, glabrous.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

nearly equaling or slightly longer than the adjacent lemmas, lanceolate to broadly lanceolate, thin, distinctly keeled, keels smooth or sparsely scabrous;

lower glumes 1-3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses glabrous or webbed, hairs usually shorter than 1/4 the lemma length, sparse;

lemmas 3-4.6 mm, lanceolate to broadly lanceolate, thin, distinctly keeled, keels and marginal veins short- to long-villous, lateral veins glabrous or sparsely softly puberulent, lateral veins obscure, intercostal regions glabrous, margins glabrous, apices acute;

paleas sparsely scabrous over the keels;

anthers (0.6)0.8-1.1(1.3) mm.

2n

= 28.

= 28, 42, 84.

Poa douglasii

Poa laxa

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ME; MT; NH; NY; OR; VT; WY; AB; BC; NL; QC
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa laxa is a low arctic to high alpine amphiatlantic species. It has been treated as a series of separate species, but the differences seem relatively minor and incomplete. Its short anthers and smoother branches usually distinguish it from P. glauca (p. 576), with which it can hybridize to form P. laxa x glauca (p. 572).

Poa laxa has four subspecies, two of which are native to the Flora region; subsp. laxa grows in central Europe; and subsp. flexuosa (Sm.) Hyl. in northwestern Europe.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Innovations primarily extravaginal; panicle branches fairly straight; calluses glabrous
subsp. banffiana
1. Innovations primarily intravaginal; panicle branches flexuous, usually at least some florets having a webbed callus
subsp. fernaldiana
Source FNA vol. 24, p. 551. FNA vol. 24, p. 570.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Oreinos
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. laxa subsp. banffiana, P. laxa subsp. fernaldiana
Name authority Nees Haenke
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