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Douglas' bluegrass, sand dune blue grass

Hartz's bluegrass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; not glaucous; densely to loosely tufted, not rhizomatous,occasionally weakly stoloniferous.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

10-33(45) cm, usually decumbent, terete;

nodes terete, 0(1) exserted.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed for 1/7 – 1/5(1/3) their length, terete, usually lustrous, bases of basal sheaths glabrous;

ligules (1.5)2-7 mm, smooth or sparsely scabrous, margins usually decurrent, apices obtuse to acuminate;

innovation blades similar in texture and shape to those of the culms;

cauline blades 2-9 cm, gradually increasing or decreasing in length upwards, 1.5-3 mm wide, folded to involute, moderately thick, soft, abaxial surfaces smooth, adaxial surfaces smooth or somewhat scabrous, usually glabrous, infrequently sparsely hispidulous, apices narrowly prow-shaped.

Basal branching

mainly intravaginal, some extravaginal.

extra- and intravaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

2.5-6(12) cm, erect, narrowly lanceolate, contracted or narrowly ovate in some bulbiferous plants, moderately congested, with 7-40 spikelets;

nodes with (1)2(4) branches;

branches 1-3 cm, erect to ascending, straight, sulcate, smooth or sparsely to moderately scabrous, with 1-10 spikelets in the distal 1/3 – 2/3.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

4.8-7.4 mm, lengths 3.5-4 times widths, lanceolate, weakly laterally compressed, sometimes bulbiferous, lustrous;

florets (2)3-5(6), normal or bulb-forming;

rachilla internodes 0.8-2 mm, smooth, sometimes sparsely hispidulous.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

mostly broadly scarious, somewhat lustrous, keels indistinct, smooth or sparsely scabrous distally;

lower glumes 3-veined;

upper glumes frequently exceeding the lowest lemmas;

calluses glabrous or with a crown of hairs, hairs to 2 mm;

lemmas (3.3)3.5-7 mm, lanceolate, usually weakly keeled, more or less evenly and somewhat loosely to densely hairy over the proximal 1/3-1/2, hairs usually longer than 0.5 mm, sparsely scabrous in the middle 1/3, smooth distally, lateral veins obscure, margins weakly inrolled, broadly scarious, glabrous, apices long-scarious, acute to shortly obtuse, often erose, often bronze-colored below the apices;

palea keels sparsely scabrous, softly puberulent at midlength, intercostal regions softly puberulent;

anthers usually all aborted late in development and 0.8-1.8 mm, infrequently well developed and 2-2.8 mm.

2n

= 28.

= 63, 70.

Poa douglasii

Poa hartzii

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NT; NU; QC; Greenland
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa hartzii grows only in the high arctic. It generally grows on open ground, on sandy or clayey soils, or on slumping slopes of old marine terraces. It carries two chloroplast genomes within its populations; one of these links it to P. secunda (p. 586) and P. ammophila (see next), the other to P. glauca (p. 576). Morphologically, it is closest to P. secunda and P. ammophila.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets bulbiferous
subsp. vrangelica
1. Spikelets not bulbiferous.
→ 2
2. Lemmas 5.5-7 mm long; anthers well developed, 2-2.8 mm long
subsp. alaskana
2. Lemmas 3.3-5.4 mm long; anthers usually aborted and shorter than 1.5 mm
subsp. hartzii
Source FNA vol. 24, p. 551. FNA vol. 24, p. 589.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Secundae > subsect. Secundae
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. hartzii subsp. alaskana, P. hartzii subsp. hartzii, P. hartzii subsp. vrangelica
Name authority Nees Gand.
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