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Douglas' bluegrass, sand dune blue grass

glaucous blue grass, timberline bluegrass, white bluegrass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; usually glaucous; densely tufted, not stoloniferous, not rhizomatous.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

5-40(80) cm, erect to spreading, straight, wiry, bases straight or slightly decumbent;

nodes terete or slightly compressed, usually 0-1 exserted, top node at 1/10-1/3 the culm length.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed for 1/10 – 1/5 their length, terete, bases of basal sheaths glabrous or sparsely minutely hairy, hairs 0.1-0.2 mm, distal sheath lengths 1.1-4 times blade lengths;

ligules 1-4(5) mm, sparsely to densely scabrous, apices obtuse to acute, minutely ciliolate;

blades 0.8-2.5 mm wide, flat or folded, thin, soft, appressed or abruptly ascending to spreading, straight, apices narrowly prow-shaped.

Basal branching

mainly intravaginal, some extravaginal.

all or mostly extravaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

1-10(20) cm, lengths 3-5 times widths at maturity, rarely racemelike with branches of irregular length, erect, narrowly lanceoloid to ovoid, contracted to somewhat open, sparse, proximal internodes shorter than 1.5(4) cm;

nodes with 2-3(5) branches;

branches erect, ascending or weakly spreading, fairly straight, short, stout, angled, angles moderately to densely scabrous, rarely only scabrous distally, glaucous;

pedicels usually shorter than the spikelets.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

3-7(9) mm, lengths 2-3 times widths, laterally compressed, rarely bulbiferous, usually glaucous;

florets 2-5, rarely bulb-forming;

rachilla internodes to 1.2 mm, smooth, muriculate, or scabrous, glabrous or sparsely to densely hispidulous or puberulent.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

subequal, narrowly to broadly lanceolate, distinctly keeled, keels smooth or sparsely scabrous, apices acute;

lower glumes 3-veined;

upper glumes 2-3.8(5.2) mm, lengths usually more than 4.1 times widths, distinctly shorter to subequal to the lowest lemmas;

calluses glabrous or webbed, webs from minute to more than 1/2 the lemma length;

lemmas 2.5-4 mm, lanceolate to broadly lanceolate, distinctly keeled, keels and marginal veins short-villous, lateral veins obscure, usually sparsely softly puberulent to short-villous, intercostal regions smooth, sometimes weakly muriculate, glabrous or puberulent, margins glabrous, apices usually partially bronze-colored, obtuse or acute;

palea keels scabrous, glabrous or softly puberulent at midlength, intercostal regions glabrous or softly puberulent;

anthers (1)1.2-2.5 mm, mature sacs 0.2 mm wide, rarely aborted late in development.

2n

= 28.

= 34, 42, 44, 47, 48, 49, 50, 56, 56, 57, 58, 60, 63, 64, 65, 70, 75, 78, ca. 100.

Poa douglasii

Poa glauca

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; NH; NM; NV; NY; OR; PA; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa glauca is a common, highly variable, circumboreal, boreal forest to alpine and high arctic species. It grows from Alaska to Greenland, south to California and New Mexico in the west, and through Canada and the northeastern United States in the east. It also grows at scattered locations in Patagonia. It generally favors dry habitats and tolerates disturbance well. It can be distinguished from P. nemoralis (p. 574) and P. interior (see previous) by its longer ligules, lower top culm node, and wider glumes and lemmas. It can be difficult to distinguish from P. laxa subsp. banffiana (p. 570). Poa glauca is often confused in herbaria with P. abbreviata subsp. pattersonii (p. 582). It differs in having primarily extravaginal branching and, usually, longer anthers. It hybridizes with P. laxa, forming P. laxa x glauca (p. 572). It is also known to hybridize with P. hartzii (p. 589), and is suspected to hybridize with P. arctica (p. 529) and P. secunda (p. 586). It is highly polyploid, and presumed to be highly apomictic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. All or some spikelets bulbiferous
var. pekulnejensis
1. Spikelets not bulbiferous.
→ 2
2. Calluses usually webbed, sometimes glabrous; lemmas glabrous or hairy between the veins
subsp. glauca
2. Calluses glabrous; lemmas hairy between the veins
subsp. rupicola
Source FNA vol. 24, p. 551. FNA vol. 24, p. 576.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Stenopoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. glauca subsp. glauca, P. glauca subsp. rupicola, P. glauca var. pekulnejensis
Name authority Nees Vahl
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