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Douglas' bluegrass, sand dune blue grass

eminent bluegrass, largeflower speargrass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; often glaucous; rhizomatous, rhizomes stout, about 2 mm thick, culms solitary.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

20-100 cm tall, about 2 mm thick, terete or weakly compressed;

nodes terete, 0-1 exserted.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed for 1/6-1/3 their length, sometimes fused by a hyaline membrane to 3/4 their length, terete, bases of some basal sheaths densely retrorsely hairy, hairs 0.1-0.2 mm, thick;

ligules 1-3.5 mm, yellow-cream to brown, truncate, erose, ciliolate;

blades (2)4-11 mm wide, flat, thick, smooth or sparsely scabrous, apices broadly prow-shaped.

Basal branching

mainly intravaginal, some extravaginal.

extravaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

8-30 cm, erect, loosely contracted, fairly congested, with 40-100+ spikelets;

branches 3-10 cm, steeply ascending, terete, smooth or sparsely scabrous, sometimes with tufts of hair at the nodes, with 5-20 spikelets.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

5-12 mm, laterally compressed;

florets 2-6;

rachilla internodes smooth, infrequently sparsely puberulent.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

lanceolate, subequal or the upper glumes to 2 mm longer than the lower glumes, sometimes exceeding the lowest lemmas, distinctly keeled, smooth, often glaucous, acute to acuminate;

lower glumes 4-9.5 mm, 1-3(5)-veined;

upper glumes 5.5-10 mm, (1)3(5)-veined;

calluses of proximal lemmas usually with a crown of hairs, hairs 1-2 mm;

lemmas 4.5-7 mm, lanceolate, 5-7-veined, distinctly keeled, thinly membranous, glabrous or the keels and marginal veins long-villous, intercostal regions glabrous or hispidulous, moderately to densely scabrous, margins usually with hairs to 0.2 mm proximally, apices acute;

palea keels scabrous;

anthers 1.7-3.2 mm.

2n

= 28.

= 28, 29+-, 42, 62.

Poa douglasii

Poa eminens

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; BC; NL; ON; QC
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa eminens grows along low arctic and boreal coasts and estuaries, in subsaline meadows and beaches. It also grows along the Asian coast from Hokkaido Island, Japan, to the Chukchi Peninsula, Russia. It hybridizes with Dupontia (see xDupoa, p. 601). Its nuclear ribosomal DNA appears to be related to an ancestor of Dupontia (p. 602) and Arctophila (p. 605); and its chloroplast DNA to P. tibetica Munro ex Stapf, an Asian member of Poa sect. Aphydris (Griseb.) Tzvelev (Gillespie & Soreng [in prep.]).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 551. FNA vol. 24, p. 598.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Arctopoa > sect. Arctopoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. rigens
Name authority Nees J. Presl
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