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Douglas' bluegrass, sand dune blue grass

Bolander's blue grass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants usually annual, rarely longer-lived; often glaucous; densely tufted, tuft bases narrow, sterile shoots few, not stoloniferous, not rhizomatous.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

20-60(70) cm, erect or geniculate at the base;

nodes terete, usually 1-3 exserted.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed for 1/2-1/4 their length, usually compressed and keeled, usually smooth, infrequently scabrous;

ligules 2.5-7 mm, smooth or scabrous, usually decurrent, obtuse to acute;

blades 1.5-5 mm wide, usually flat, rarely folded, lax, soft, smooth or sparsely scabrous, margins scabrous, apices broadly prow-shaped, cauline blades 3-15 cm, flag leaf blades 1-4 cm.

Basal branching

mainly intravaginal, some extravaginal.

both intra- and extravaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

(5)10-15(25) cm long, 1/4-1/2 the plant height, usually erect, infrequently slightly nodding, usually eventually open, sometimes interrupted, sparse, with 1-3(5) branches per node;

branches initially erect and straight, usually some eventually spreading or reflexed, smooth or sparsely to moderately scabrous.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

(3)4-7 mm, laterally compressed;

florets 2-3(4);

rachilla internodes usually 1-1.2+ mm, smooth or sparsely scabrous, glabrous.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

unequal, distinctly shorter than the adjacent lemmas, distinctly keeled, keels smooth or sparsely scabrous;

lower glumes 1-3-veined, 1/2 - 2/3 the length of the upper glume, 1/2 - 2/3 the length of the lowest lemmas;

upper glumes shorter than or subequal to the lowest lemmas;

calluses of some or all florets sparsely webbed;

lemmas 2.5-4 mm, lanceolate to narrowly lanceolate, distinctly keeled, smooth or scabrous throughout, glabrous, lateral veins obscure to moderately prominent, apices narrowly acute, usually anthocyanic near the tip;

palea keels sparsely scabrous;

anthers 3, 0.5-1(1.8) mm.

2n

= 28.

= 28.

Poa douglasii

Poa bolanderi

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; NV; OR; UT; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa bolanderi grows mainly in pine to fir forest openings of mountain slopes in the western United States, from Washington to California and Utah. It differs from P. howellii (see below) in having smooth to scabrous, rather than puberulent, lemmas; it also grows at higher elevations, mostly at 1500-3000 m.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 551. FNA vol. 24, p. 533.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. howellii var. chandleri
Name authority Nees Vasey
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