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Douglas' bluegrass, sand dune blue grass

Texas bluegrass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; loosely tufted, rhizomatous, rhizomes slender.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

20-85 cm, erect, terete or weakly compressed;

nodes terete, 0-1 exserted.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed firmly for 1/7-1/3 their length, sometimes for a longer distance by a hyaline membrane, terete, smooth, glabrous, bases of basal sheaths glabrous;

ligules 1-4 mm, smooth or scabrous;

innovation blades 10-35 cm long, 1-3.5 mm wide;

cauline blades 2-25 cm long, 1.5-4.5 mm wide, flat or folded, lax, both surfaces smooth or sparsely finely scabrous, glabrous, apices narrowly to broadly prow-shaped.

Basal branching

mainly intravaginal, some extravaginal.

intra- and extravaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

3-12(18) cm, erect, narrowly cylindrical, often interrupted or lobed, congested, with (70)100-200 spikelets;

nodes with (2)3-7(9) branches;

branches 1-3(5) cm, erect to slightly ascending, terete or weakly angled, sparsely to densely coarsely scabrous, with 8-30 spikelets.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

4-8(10) mm, sexually dimorphic, laterally compressed, pistillate spikelets larger, with fewer florets and more pubescence than the staminate spikelets;

florets 2-10;

rachilla internodes smooth.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

unequal, distinctly keeled, keels and lateral veins scabrous;

lower glumes 1-3-veined.

Pistillate florets

calluses copiously 3-webbed, hairs 4-10 mm, mostly silky, plicate;

lemmas 4.2-6.4 mm, lanceolate, 5-7 veined, distinctly keeled, glabrous, or the keels and marginal veins, sometimes also the lateral veins, densely long-villous, margins glabrous, apices acute;

palea keels scabrous, glabrous or sometimes softly puberulent to long-villous at midlength.

Staminate florets

calluses glabrous or sparsely dorsally webbed, hairs plicate, rarely with additional webs under the marginal veins;

lemmas 3.5-5 mm, lanceolate, distinctly keeled, keels and marginal veins sparsely short- to long-villous, margins glabrous, apices acute;

palea keels scabrous, glabrous or softly puberulent to long-villous at midlength;

anthers vestigial (0.1-0.2 mm) or 1.6-2.7 mm.

2n

= 28.

= 42, ca. 54, 56, ca. 63, 84.

Poa douglasii

Poa arachnifera

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; IL; KS; LA; MS; NC; NM; OK; SC; TN; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa arachnifera grows on moist, sandy to rich, black bottomlands of the southern Great Plains. At one time it was cultivated for winter pasture in the southeastern United States. It is strictly dioecious, with a 1:1 ratio of staminate to pistillate plants among herbarium samples. The variable and high chromosome numbers suggest it may be apomictic, but the occurrence of equal numbers of staminate and pistillate individuals in populations seems to suggest that reproduction is primarily sexual. It is the only non-South American species in the section. Its closest relatives appear to be P. bonariensis (Lam.) Kunth and P. lanuginosa Poir.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 551. FNA vol. 24.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Dioicopoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Name authority Nees Torr.
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