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Douglas' bluegrass, sand dune blue grass

alpine blue grass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; not glaucous; densely cespitose, not rhizomatous, not stoloniferous.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

10-40 cm.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

Basal branching

mainly intravaginal, some extravaginal.

intravaginal.

Leaves

mostly basal;

sheaths closed for 1/8 – 2/7 their length, terete, basal sheaths persistent, overlapping, bases usually not swollen;

ligules of innovations 1-2(3) mm, those of the upper cauline leaves to 4(5) mm, milky white, smooth, glabrous, obtuse;

blades of innovations widely spreading, persisting through the season, blades of cauline leaves 1-5(12) cm long, 2-4.5 mm wide, flat, moderately thick, soft, straight, smooth or the margins sparsely scabrous, apices broadly prow-shaped, blades of upper cauline leaves much reduced in length.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

2-6(8) cm, erect, ovoid to pyramidal, open or loosely contracted at maturity, fairly congested;

nodes with 1-2 branches, lowest internodes 0.6-1(1.5) cm;

branches 1-3(4) cm, ascending to spreading, straight, terete, usually smooth or sparsely scabrous, rarely moderately densely scabrous;

pedicels divaricate, shorter than the spikelets.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

3.9-6.2 mm, ovate, lengths 1.5-2.5 times widths, laterally compressed, plump, sometimes bulbiferous;

florets 3-7, usually normal;

rachilla internodes 0.5-0.8 mm, smooth, glabrous or sparsely softly puberulent to short-villous.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

broadly lanceolate to narrowly ovate, keeled, keels sparsely scabrous;

lower glumes 3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses glabrous;

lemmas 3-5 mm, broadly lanceolate, keeled, keels and marginal veins short- to long-villous, lateral veins moderately prominent, intercostal regions sparsely to moderately short-villous, apices acute;

palea keels softly puberulent to short-villous over most of their length, apices scabrous;

anthers 1.3-2.3 mm.

2n

= 28.

= 22, 23, 24,25, 26, 27, 28, 28+11, 30, 31, 32, 32+1, 33, 34, 35, 36, 37, 39, 40+1, 41, 42, ca. 43, 44, 46, ca. 48, 56.

Poa douglasii

Poa alpina

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CO; ID; MI; MT; NM; NV; OR; UT; WA; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa alpina is a fairly common circumboreal forest species of subalpine to arctic habitats, extending south in the Rocky Mountains to Utah and Colorado in the west, and to the northern Great Lakes region in the east. It often grows in disturbed ground and is calciphilic. Poa xgaspensis (p. 601) is a natural hybrid which seems to be between P. alpina and P. pratensis subsp. alpigena (p. 525); it differs from P. alpina in its extravaginal branching, rhizomatous habit, and webbed calluses. The range of chromosome numbers suggests that P. alpina is predominantly apomictic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets not bulbiferous
subsp. alpina
1. Some or all spikelets bulbiferous
subsp. vivipara
Source FNA vol. 24, p. 551. FNA vol. 24.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Alpinae
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. alpina subsp. alpina, P. alpina subsp. vivipara
Name authority Nees L.
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