Poa cuspidata |
Poaceae subfam. pooideae |
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early bluegrass |
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Habit | Plants perennial; loosely tufted or with solitary shoots, shortly rhizomatous. | Plants annual or perennial; sometimes matlike, sometimes cespitose, sometimes stoloniferous, sometimes rhizomatous. | ||||||||||||||||||||||||||||||||||||
Culms | 15-60 cm, erect or the bases decumbent, not branching above the base, terete or weakly compressed; nodes terete, 0-1 exserted. |
usually hollow, sometimes solid. |
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Sheaths | closed for about 1/2 their length, slightly compressed, distinctly keeled, glabrous, bases of basal sheaths glabrous, distal sheath lengths 4-60 times blade lengths; collars of proximal leaves usually retrorsely scabrous or pubescent distally and about the throat; ligules 0.5-4 mm, smooth or scabrous, apices truncate to acute; innovation blades similar to the cauline blades; cauline blades 1-4 mm wide, usually flat, sometimes slightly folded, smooth or sparsely scabrous, primarily over the veins, apices broadly prow-shaped, blades steeply reduced in length distally, flag leaf blades 0.2-3(6) cm. |
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Basal branching | mainly extravaginal. |
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Leaves | distichous; sheaths usually open to the base, varying to closed for nearly their full length; auricles present or absent; abaxial ligules absent; adaxial ligules scarious or membranous, sometimes puberulent or scabridulous, usually not ciliate, cilia sometimes shorter than the base; pseudopetioles rarely present; blades usually linear, sometimes broadly so, venation parallel; cross sections non-Kranz, mesophyll nonradiate, adaxial palisade layer absent, fusoid and arm cells usually absent; midribs usually simple; adaxial bulliform cells present; stomates with parallel-sided subsidiary cells; epidermes usually lacking bicellular microhairs, sometimes with unicellular microhairs, papillae usually absent, when present, rarely more than 1 per cell. |
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Panicles | 5-15 cm, erect or lax, pyramidal, open, sparse, with 20-80 spikelets, proximal internodes usually 3+ cm; nodes usually with 2 branches; branches (2)3-7(10) cm, spreading to reflexed, straight, angled, angles scabrous, with 2-8(10) spikelets. |
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Inflorescences | usually terminal, panicles, spikes, or racemes, usually ebracteate; disarticulation usually below the florets, sometimes below the glumes, at the rachis nodes, or at the inflorescence bases. |
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Spikelets | 5-8 mm, lengths to 3.5 times widths, laterally compressed, not sexually dimorphic; florets 2-5; rachilla internodes smooth. |
usually bisexual, infrequently unisexual or mixed, usually laterally compressed or not compressed, occasionally dorsally compressed, with 1-30 sexual florets, distal floret(s) often reduced, infrequently spikelets with 1-2 reduced or staminate basal florets and a single terminal sexual floret. |
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Glumes | narrowly lanceolate to lanceolate, distinctly keeled; lower glumes 1-3-veined; calluses webbed, hairs over 1/3 the lemma length; lemmas 3-6 mm, lanceolate, distinctly keeled, keels and marginal veins sparsely short- to long-villous, lateral veins moderately prominent, intercostal regions glabrous or the upper florets in the spikelets softly puberulent, margins glabrous, apices acute; palea keels scabrous, softly puberulent at midlength; anthers vestigial (0.1-0.2 mm) or 2-3.5 mm. |
usually 2, upper or lower glumes sometimes absent, rarely both glumes absent; lemmas without uncinate hairs, awned or not, awns single, basal to apical; paleas usually well-developed, sometimes reduced or absent; lodicules 2(3), usually lanceolate and broadly membranous distally, rarely truncate and fleshy, usually not veined or obscurely veined, sometimes distinctly veined, sometimes ciliate; anthers (1, 2)3; ovaries glabrous or sometimes hairy distally, sometimes with an apical appendage; haustorial synergids absent; styles (1)2 (-4), bases close together, sometimes fused. |
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Caryopses | hila linear, elliptic, ovate, or punctate; endosperm usually hard, sometimes soft or liquid, with or without lipids, starch grains compound or simple; embryos less than 1/2 the length of the caryopses; epiblasts usually present; scutellar cleft usually absent; mesocotyl internode usually absent; embryonic leaf margins overlapping, x = 7, 10. |
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2n | = 28. |
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Poa cuspidata |
Poaceae subfam. pooideae |
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Distribution |
AL; DC; DE; GA; IN; KY; LA; MD; NC; NJ; NY; OH; PA; SC; TN; VA; WV
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Discussion | Poa cuspidata is a common species of forest openings in the Appalachian Mountains. It is an eastern counterpart of P. arnowiae (see previous), P. tracyi (p. 543), and P. nervosa (see next). Like those species, it is sequentially gynomonoecious. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The subfamily Pooideae includes approximately 3300 species, making it the largest subfamily in the Poaceae. It reaches its greatest diversity in cool temperate and boreal regions, extending across the tropics only in high mountains. The circumscription and relationships of tribes within the Pooideae are unsettled (see, for example, Catalan et al. 1997, 2004; Soreng and Davis 1998). In this flora, some previously recognized tribes have been combined with the Poeae. Recognition of some of these as subtribes is well supported; among these is the Hainardieae Greuter (which, at the subtribal level, is called the Parapholiinae Caro). Members of other traditional tribal groupings, such as the Aveneae Dumort., appear to be widely dispersed within the Poeae sensu lato. Further work will probably support the division of the expanded Poeae into additional tribes; there is as yet no clear indication as to what the boundaries of such tribes should be. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 545. | FNA vol. 24, p. 57. | ||||||||||||||||||||||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa | Poaceae | ||||||||||||||||||||||||||||||||||||
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Name authority | Nutt. | Benth. | ||||||||||||||||||||||||||||||||||||
Web links |