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early bluegrass

Hooker's bluegrass, Wheeler's blue grass

Habit Plants perennial; loosely tufted or with solitary shoots, shortly rhizomatous. Plants perennial; densely to loosely tufted or with solitary shoots, shortly rhizomatous.
Culms

15-60 cm, erect or the bases decumbent, not branching above the base, terete or weakly compressed;

nodes terete, 0-1 exserted.

35-80 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 1-2 exserted.

Sheaths

closed for about 1/2 their length, slightly compressed, distinctly keeled, glabrous, bases of basal sheaths glabrous, distal sheath lengths 4-60 times blade lengths;

collars of proximal leaves usually retrorsely scabrous or pubescent distally and about the throat;

ligules 0.5-4 mm, smooth or scabrous, apices truncate to acute;

innovation blades similar to the cauline blades;

cauline blades 1-4 mm wide, usually flat, sometimes slightly folded, smooth or sparsely scabrous, primarily over the veins, apices broadly prow-shaped, blades steeply reduced in length distally, flag leaf blades 0.2-3(6) cm.

closed for 1/3-1/4 their length, terete to slightly compressed, at least some proximal sheaths densely retrorsely scabrous, hispidulous, or softly puberulent for the upper 1/4 of their length, bases of basal sheaths glabrous, distal sheath lengths (1.4)1.7-4.6(6.2) times blade lengths;

collars of proximal leaves glabrous or with hairs the same length as those of their sheaths;

ligules 0.5-2 mm, smooth or scabrous, sometimes puberulent, truncate, those of the lower culm and innovation leaves 0.5-1.5 mm, abaxial surfaces scabrous to softly puberulent, truncate;

innovation blades folded or involute, infrequently flat, moderately thick, soft, adaxial surfaces usually densely scabrous to hispidulous;

cauline blades 2-3.5 mm wide, flat or folded, smooth or sparsely scabrous, glabrous or hispidulous, apices narrowly to broadly prow-shaped, blades gradually reduced distally or the middle blades longest, flag leaf blades 1-10 cm long.

Basal branching

mainly extravaginal.

mainly extra-vaginal.

Panicles

5-15 cm, erect or lax, pyramidal, open, sparse, with 20-80 spikelets, proximal internodes usually 3+ cm;

nodes usually with 2 branches;

branches (2)3-7(10) cm, spreading to reflexed, straight, angled, angles scabrous, with 2-8(10) spikelets.

5-12(18) cm, erect or nodding, ovoid to pyramidal, loosely contracted to open, with 20-70 spikelets, proximal internodes usually shorter than 3.5 cm;

nodes with 2-5 branches;

branches (1)1.7-6.5 cm, ascending to spreading or reflexed, lax, terete or weakly angled, sparsely to moderately scabrous, with 2-8(12) spikelets.

Spikelets

5-8 mm, lengths to 3.5 times widths, laterally compressed, not sexually dimorphic;

florets 2-5;

rachilla internodes smooth.

5.5-10 mm, lengths to 3.5 times widths, laterally compressed, not sexually dimorphic;

florets 2-7;

rachilla internodes smooth or scabrous, glabrous or sparsely to densely hispidulous.

Glumes

narrowly lanceolate to lanceolate, distinctly keeled;

lower glumes 1-3-veined;

calluses webbed, hairs over 1/3 the lemma length;

lemmas 3-6 mm, lanceolate, distinctly keeled, keels and marginal veins sparsely short- to long-villous, lateral veins moderately prominent, intercostal regions glabrous or the upper florets in the spikelets softly puberulent, margins glabrous, apices acute;

palea keels scabrous, softly puberulent at midlength;

anthers vestigial (0.1-0.2 mm) or 2-3.5 mm.

1/4 - 2/3(3/4) as long as the adjacent lemmas, lanceolate, distinctly keeled;

lower glumes 1-3-veined, 1/4-1/2 as long as the adjacent lemmas;

calluses glabrous;

lemmas 3-6 mm, lanceolate, distinctly keeled, keels and marginal veins glabrous or softly puberulent to short-villous, intercostal regions glabrous or hispidulous, infrequently puberulent, smooth or finely scabrous, lateral veins obscure to moderately prominent, margins glabrous, apices acute;

palea keels scabrous, intercostal regions glabrous;

anthers usually vestigial (0.1-0.2 mm) or aborted late in development and up to 2 mm, rarely normal.

2n

= 28.

= 56, 61, 62, 63, 64, 66, 61, 70, ca. 74, 75, 79, 80, 81, 87, 89, 90, 91.

Poa cuspidata

Poa wheeleri

Distribution
from FNA
AL; DC; DE; GA; IN; KY; LA; MD; NC; NJ; NY; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; MB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa cuspidata is a common species of forest openings in the Appalachian Mountains. It is an eastern counterpart of P. arnowiae (see previous), P. tracyi (p. 543), and P. nervosa (see next). Like those species, it is sequentially gynomonoecious.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa wheeleri is common at mid- to high elevations, generally on the east side of the coastal mountains from British Columbia to California, and from Manitoba to New Mexico. It generally grows in submesic coniferous forests to subalpine habitats. Most plants have densely retrorsely pubescent or scabrous sheaths, involute innovation blades that are pubescent adaxially, and pistillate florets.

Poa wheeleri, a high polyploid apomictic species, probably arose from hybridization between P. cusickii (p. 559) and another member of the Poa nervosa complex. It resembles P. rhizomata (see next) and P. chambersii (p. 548) more than P. nervosa sensu stricto (see previous). It differs from P. chambersii in having at least some proximal sheaths that are densely retrorsely scabrous or pubescent (sometimes obscurely so), and folded or involute innovation blades that are scabrous to hispidulous on the adaxial surfaces. For a comparison with P. nervosa, see p. 545. Natural hybrids have been found between P. wheeleri and P. pratensis (p. 522).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 545. FNA vol. 24, p. 546.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. nervosa var. wheeleri, P. curta
Name authority Nutt. Vasey
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