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early bluegrass

ocean bluff blue grass, San Francisco bluegrass, sea-bluff bluegrass

Habit Plants perennial; loosely tufted or with solitary shoots, shortly rhizomatous. Plants perennial; frequently glaucous; densely tufted, not stoloniferous, not rhizomatous.
Culms

15-60 cm, erect or the bases decumbent, not branching above the base, terete or weakly compressed;

nodes terete, 0-1 exserted.

5-40 cm, erect or ascending, frequently decumbent, terete, with 0-2 exserted nodes.

Sheaths

closed for about 1/2 their length, slightly compressed, distinctly keeled, glabrous, bases of basal sheaths glabrous, distal sheath lengths 4-60 times blade lengths;

collars of proximal leaves usually retrorsely scabrous or pubescent distally and about the throat;

ligules 0.5-4 mm, smooth or scabrous, apices truncate to acute;

innovation blades similar to the cauline blades;

cauline blades 1-4 mm wide, usually flat, sometimes slightly folded, smooth or sparsely scabrous, primarily over the veins, apices broadly prow-shaped, blades steeply reduced in length distally, flag leaf blades 0.2-3(6) cm.

closed for 1/10(1/5) their length, terete, smooth, glabrous, bases of basal sheaths glabrous, distal sheath lengths 1-4 times widths;

ligules 2-6 mm, smooth or sparsely scabrous, obtuse to acute;

innovation blades usually 1-1.5 mm wide, thin, soon withering, and distinctly narrower than the cauline blades, infrequently wider, flat, and a bit fleshy as in the cauline blades, or involute;

cauline blades gradually reduced in length distally, 2-5 mm wide, flat or folded, soft, thin and soon withering or moderately thick and somewhat fleshy and retaining their form, smooth, apices narrowly to broadly prow-shaped.

Basal branching

mainly extravaginal.

all or mainly intravaginal.

Panicles

5-15 cm, erect or lax, pyramidal, open, sparse, with 20-80 spikelets, proximal internodes usually 3+ cm;

nodes usually with 2 branches;

branches (2)3-7(10) cm, spreading to reflexed, straight, angled, angles scabrous, with 2-8(10) spikelets.

3-7 cm, erect, nearly cylindrical, contracted, congested, with (20)30-80(120) spikelets;

nodes with 3-7 branches;

branches 0.5-1.5(4.5) cm, erect, usually angled, infrequently terete or sulcate, angles usually moderately to densely scabrous, infrequently smooth with dense papillae and weak-angled, with 2-10 spikelets in the proximal 2/3;

pedicels shorter than the spikelets.

Spikelets

5-8 mm, lengths to 3.5 times widths, laterally compressed, not sexually dimorphic;

florets 2-5;

rachilla internodes smooth.

4.5-7 mm, lengths to 3.5 times widths, lanceolate to narrowly ovate, drab;

florets 3-5;

rachilla internodes usually shorter than 1 mm, smooth.

Glumes

narrowly lanceolate to lanceolate, distinctly keeled;

lower glumes 1-3-veined;

calluses webbed, hairs over 1/3 the lemma length;

lemmas 3-6 mm, lanceolate, distinctly keeled, keels and marginal veins sparsely short- to long-villous, lateral veins moderately prominent, intercostal regions glabrous or the upper florets in the spikelets softly puberulent, margins glabrous, apices acute;

palea keels scabrous, softly puberulent at midlength;

anthers vestigial (0.1-0.2 mm) or 2-3.5 mm.

lanceolate, slightly unequal, distinctly keeled, keels papillate or scabrous;

lower glumes 3-veined;

calluses glabrous or with a crown of hairs, hairs 0.1-0.2 mm;

lemmas 3-4.5 mm, lanceolate, distinctly keeled, glabrous or the keels and marginal veins short-villous to midlength, intercostal regions sparsely puberulent near the base, margins glabrous, apices acute;

palea keels scabrous, sometimes softly puberulent at midlength;

anthers 1.5-3 mm.

2n

= 28.

= 42, 84.

Poa cuspidata

Poa unilateralis

Distribution
from FNA
AL; DC; DE; GA; IN; KY; LA; MD; NC; NJ; NY; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa cuspidata is a common species of forest openings in the Appalachian Mountains. It is an eastern counterpart of P. arnowiae (see previous), P. tracyi (p. 543), and P. nervosa (see next). Like those species, it is sequentially gynomonoecious.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa unilateralis grows on grassy bluffs and cliffs near the Pacific coast of North America, from Washington to California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lemmas villous on the keels and marginal veins for more than 1/3 the length of the lemmas; blades usually involute, the cauline and innovation blades similar
subsp. pachypholis
1. Lemmas glabrous or the keels and marginal veins villous or puberulent for less than 1/5 the length of the lemmas; blades flat or folded, the cauline blades sometimes wider and thicker than the innovation blades
subsp. unilateralis
Source FNA vol. 24, p. 545. FNA vol. 24, p. 594.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Secundae > subsect. Halophytae
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. unilateralis subsp. pachypholis, P. unilateralis subsp. unilateralis
Name authority Nutt. Scribn.
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