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early bluegrass

Howell's blue grass

Habit Plants perennial; loosely tufted or with solitary shoots, shortly rhizomatous. Plants usually annual, rarely longer-lived; densely tufted, tuft bases narrow; not stoloniferous, not rhizomatous.
Culms

15-60 cm, erect or the bases decumbent, not branching above the base, terete or weakly compressed;

nodes terete, 0-1 exserted.

(10) 25-80(120) cm tall, 0.4-1.75 mm thick, usually erect;

nodes terete, usually 1-2 exserted.

Sheaths

closed for about 1/2 their length, slightly compressed, distinctly keeled, glabrous, bases of basal sheaths glabrous, distal sheath lengths 4-60 times blade lengths;

collars of proximal leaves usually retrorsely scabrous or pubescent distally and about the throat;

ligules 0.5-4 mm, smooth or scabrous, apices truncate to acute;

innovation blades similar to the cauline blades;

cauline blades 1-4 mm wide, usually flat, sometimes slightly folded, smooth or sparsely scabrous, primarily over the veins, apices broadly prow-shaped, blades steeply reduced in length distally, flag leaf blades 0.2-3(6) cm.

closed for 1/2 - 7/8 their length, usually weakly compressed and keeled, usually scabrous, rarely smooth;

ligules 1.5-5(10) mm, smooth or scabrous, acute;

blades 1-7(10) mm wide, flat, lax, soft, finely scabrous, apices narrowly prow-shaped, cauline blades 2-10 cm.

Basal branching

mainly extravaginal.

intravaginal.

Panicles

5-15 cm, erect or lax, pyramidal, open, sparse, with 20-80 spikelets, proximal internodes usually 3+ cm;

nodes usually with 2 branches;

branches (2)3-7(10) cm, spreading to reflexed, straight, angled, angles scabrous, with 2-8(10) spikelets.

10-25(30) cm, erect, eventually open, with (1)3-5(7) branches per node;

branches eventually spreading or reflexed, fairly straight, angled, angles usually moderately to densely scabrous, rarely sparsely scabrous.

Spikelets

5-8 mm, lengths to 3.5 times widths, laterally compressed, not sexually dimorphic;

florets 2-5;

rachilla internodes smooth.

(2)4-6 mm, laterally compressed, with 2-5 florets;

rachilla internodes about 1 mm, smooth, usually softly puberulent, infrequently glabrous.

Glumes

narrowly lanceolate to lanceolate, distinctly keeled;

lower glumes 1-3-veined;

calluses webbed, hairs over 1/3 the lemma length;

lemmas 3-6 mm, lanceolate, distinctly keeled, keels and marginal veins sparsely short- to long-villous, lateral veins moderately prominent, intercostal regions glabrous or the upper florets in the spikelets softly puberulent, margins glabrous, apices acute;

palea keels scabrous, softly puberulent at midlength;

anthers vestigial (0.1-0.2 mm) or 2-3.5 mm.

slightly unequal, lanceolate, distinctly keeled, keels and sometimes the lateral veins sparsely to moderately scabrous;

lower glumes 1-3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses of some or all florets sparsely webbed;

lemmas 2.5-3.5 mm, lanceolate to narrowly lanceolate, distinctly keeled, crisply puberulent proximally, hairs evenly distributed, finely scabrous distally, lateral veins obscure to prominent, margins narrowly hyaline, glabrous, apices narrowly acute, infrequently anthocyanic;

palea keels sparsely scabrous, glabrous or softly puberulent at midlength, intercostal regions usually softly puberulent;

anthers 3, 0.2-1 mm.

2n

= 28.

= unknown.

Poa cuspidata

Poa howellii

Distribution
from FNA
AL; DC; DE; GA; IN; KY; LA; MD; NC; NJ; NY; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa cuspidata is a common species of forest openings in the Appalachian Mountains. It is an eastern counterpart of P. arnowiae (see previous), P. tracyi (p. 543), and P. nervosa (see next). Like those species, it is sequentially gynomonoecious.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa howellii grows primarily on rocky banks and wooded slopes, from the coastal ranges of southern British Columbia to southern California. It differs from P. bolanderi (see above) in having puberulent, rather than smooth or scabrous, lemmas, and in growing at lower elevations, mostly from near sea level to 1000 m.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 545. FNA vol. 24, p. 534.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. bolanderi var. howellii
Name authority Nutt. Vasey & Scribn.
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