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Cusick's bluegrass

supine bluegrass

Habit Plants perennial; usually densely tufted, rarely moderately densely tufted, usually neither rhizomatous nor stoloniferous, infrequently short-rhizomatous or stoloniferous, rarely with distinct rhizomes. Plants perennial; stoloniferous, loosely tufted.
Culms

10-60(70) cm tall, 0.5-1.8 mm thick, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-2 exserted.

8-12(20) cm, slender, bases decumbent, terete or weakly compressed;

nodes terete, 1 exserted.

Sheaths

closed for 1/4-3/4 their length, terete, smooth or scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths 1.6-10 times blade lengths;

collars smooth or scabrous, glabrous;

ligules of cauline leaves 1-3(6) mm, smooth or scabrous, truncate to acute, ligules of the innovation leaves 0.2-0.5(2.5) mm, scabrous, usually truncate;

innovation blades sometimes distinctly different from the cauline blades, 0.5-2 mm wide, involute, moderately thick, moderately firm, adaxial surfaces usually densely scabrous or hispidulous to softly puberulent, infrequently nearly smooth and glabrous;

cauline blades subequal or the midcauline blades longest or the blades gradually reduced in length distally, 0.5-3 mm wide, flat, folded, or involute, usually thin, usually withering, abaxial surfaces smooth or scabrous, apices narrowly to broadly prow-shaped, flag leaf blades 0.5-5(6) cm.

closed for 1/4-1/3 their length, terete, smooth, glabrous, bases of basal sheaths glabrous, distal sheath lengths 2-4 times blade lengths;

collars smooth, glabrous;

ligules 0.6-1 mm, smooth, glabrous, truncate;

blades 2-3 mm wide, flat, thin, soft, smooth, apices broadly prow-shaped, cauline blades subequal.

Basal branching

intravaginal or intra- and extravaginal.

intravaginal.

Panicles

2-10(12) cm, usually erect, contracted or loosely contracted, narrowly lanceoloid to ovoid, congested or moderately congested, with 10-100 spikelets and 1-3(5) branches per node;

branches 0.5-4(5) cm, erect or steeply ascending, fairly straight, slender to stout, terete to angled, smooth or scabrous, with 1-15 spikelets.

2.5-5 cm, lengths 1-2 times widths, erect, loosely contracted or open, ovoid to pyramidal, sparse, with 10-25(30) spikelets and 1-2 branches per node;

branches 1-3 cm, spreading to reflexed, straight, terete, smooth or sparsely scabrous, with 2-5(8) spikelets.

Spikelets

(3)4-10 mm, lengths to 3 times widths, broadly lanceolate to narrowly ovate, laterally compressed, not sexually dimorphic;

florets 2-6;

rachilla internodes 0.5-1.2 mm, smooth or scabrous.

4-6 mm, laterally compressed;

florets 3-7;

rachilla internodes smooth, glabrous, more or less concealed, distal internode less than 1/2 the length of the distal lemma.

Glumes

lanceolate, distinctly keeled;

lower glumes 3-veined, distinctly shorter than the lowest lemmas;

calluses glabrous or diffusely webbed, hairs less than 1/4 the lemma length;

lemmas (3)4-7 mm, lanceolate to broadly lanceolate, distinctly keeled, membranous to thinly membranous, smooth or sparsely to densely scabrous, glabrous or the keels and/or marginal veins puberulent proximally, lateral veins obscure to prominent, margins glabrous, apices acute;

palea keels scabrous, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm), aborted late in development, or 2-3.5 mm.

distinctly keeled, keels smooth;

lower glumes 1-veined;

calluses glabrous;

lemmas 1.7-4 mm, lanceolate, distinctly keeled, smooth throughout, proximal lemmas glabrous throughout or the keels and marginal veins sparsely short-villous, distal lemmas glabrous or the keels and marginal veins short-villous to near the apices, lateral veins moderately prominent, intercostal regions glabrous, margins smooth, glabrous, apices obtuse to acute;

palea keels smooth, sometimes sparsely softly puberulent to short-villous;

anthers (1.25)1.5-2.5 mm, cylindrical prior to dehiscence, those of the upper 1-2 florets commonly vestigial.

2n

= 28, 28+11, 56, 56+11, 59, ca. 70.

= 14.

Poa cusickii

Poa supina

Distribution
from FNA
CA; CO; ID; MT; ND; NV; OR; UT; WA; WY; AB; BC; MB; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa cusickii grows in rich meadows in sagebrush scrub to rocky alpine slopes, from the southwestern Yukon Territory to Manitoba and North Dakota, south to central California and eastern Colorado. It is gynodioecious or dioecious.

Sexually reproducing plants of Poa cusickii subspp. cusickii and pallida grow in different geographic areas, but pistillate plants of these two subspecies have overlapping ranges. Only pistillate plants are known in Poa cusickii subspp. epilis and purpurascens. All the alpine plants studied were pistillate.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa supina is native to boreal to alpine regions of Eurasia. Beginning in the 1990s, the cultivar 'Supernova' has been introduced for seeding in wet to moist, cool, shady areas subject to heavy traffic. It has been tested in both Canada and the United States, and is expected to gradually escape cultivation, probably becoming established throughout the cool-temperate portion of the Flora region. Its current distribution is not known. Poa supina differs from P. annua (p. 519), of which is thought to be one of the parents, in having longer anthers and a more stoloniferous habit, as well as in being diploid. It is gynomonoecious.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Panicle branches smooth or slightly scabrous, or the basal blades more than 1.5 mm wide and flat or folded; cauline blades more than 1.5 mm wide, often flat; some basal branching extravaginal; lemmas and calluses sometimes sparsely puberulent.
→ 2
2. Lemmas usually glabrous, rarely plants from the Rocky Mountains with puberulent keels and marginal veins; calluses glabrous; panicles erect, usually contracted; branches smooth to slightly scabrous
subsp. epilis
2. Lemmas rarely completely glabrous, at least some florets v/ith sparsely puberulent keels, the marginal veins glabrous or puberulent; calluses frequendy with a sparse, short web; panicles somewhat lax and loosely contracted; branches smooth or sparsely to moderately scabrous
subsp. purpurascens
1. Panicle branches moderately to strongly scabrous; basal and cauline blades usually less than 1.5 mm wide, involute, rarely flat or folded; basal branching intravaginal; lemmas and calluses glabrous.
→ 3
3. Panicle branches longer than 1.7 cm in at least some panicles; panicles open or contracted
subsp. cusickii
3. Panicle branches up to 1.7 cm long, stout; panicles contracted
subsp. pallida
Source FNA vol. 24, p. 559. FNA vol. 24, p. 521.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Epiles Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Micrantherae
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. cusickii subsp. cusickii, P. cusickii subsp. epilis, P. cusickii subsp. pallida, P. cusickii subsp. purpurascens
Name authority Vasey Schrad.
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