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Cusick's bluegrass

Eurasian woodland bluegrass, forest blue grass, interior bluegrass (ssp. interior), wood blue grass, woodland bluegrass, woods bluegrass (ssp. nemoralis)

Habit Plants perennial; usually densely tufted, rarely moderately densely tufted, usually neither rhizomatous nor stoloniferous, infrequently short-rhizomatous or stoloniferous, rarely with distinct rhizomes. Plants perennial; green or glaucous; densely tufted, not stoloniferous, not rhizomatous.
Culms

10-60(70) cm tall, 0.5-1.8 mm thick, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-2 exserted.

30-80 cm, mostly erect, smooth below the panicles;

nodes slightly compressed, 2-5 exserted, top node at 1/2 - 3/4 the culm length.

Sheaths

closed for 1/4-3/4 their length, terete, smooth or scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths 1.6-10 times blade lengths;

collars smooth or scabrous, glabrous;

ligules of cauline leaves 1-3(6) mm, smooth or scabrous, truncate to acute, ligules of the innovation leaves 0.2-0.5(2.5) mm, scabrous, usually truncate;

innovation blades sometimes distinctly different from the cauline blades, 0.5-2 mm wide, involute, moderately thick, moderately firm, adaxial surfaces usually densely scabrous or hispidulous to softly puberulent, infrequently nearly smooth and glabrous;

cauline blades subequal or the midcauline blades longest or the blades gradually reduced in length distally, 0.5-3 mm wide, flat, folded, or involute, usually thin, usually withering, abaxial surfaces smooth or scabrous, apices narrowly to broadly prow-shaped, flag leaf blades 0.5-5(6) cm.

closed for 1/10 – 1/5 their length, terete, bases of basal sheaths glabrous, distal sheath lengths 0.45-1 (1.1) times blade lengths;

ligules 0.2-0.8(1) mm, sparsely to densely scabrous, apices truncate, minutely ciliolate;

blades 0.8-3 mm wide, mostly flat, appressed, abruptly ascending to spreading, straight or somewhat lax, apices narrowly prow-shaped.

Basal branching

intravaginal or intra- and extravaginal.

all or mostly extravaginal.

Panicles

2-10(12) cm, usually erect, contracted or loosely contracted, narrowly lanceoloid to ovoid, congested or moderately congested, with 10-100 spikelets and 1-3(5) branches per node;

branches 0.5-4(5) cm, erect or steeply ascending, fairly straight, slender to stout, terete to angled, smooth or scabrous, with 1-15 spikelets.

7-16(20) cm, lengths usually 2.5-4 times widths at maturity, usually erect, lax in shade forms, narrowly lanceoloid to ovoid, slightly to moderately congested;

nodes with 2-5 branches;

branches ascending to widely spreading, fairly straight, slender to moderately stout, angled, angles moderately to densely scabrous.

Spikelets

(3)4-10 mm, lengths to 3 times widths, broadly lanceolate to narrowly ovate, laterally compressed, not sexually dimorphic;

florets 2-6;

rachilla internodes 0.5-1.2 mm, smooth or scabrous.

3-8 mm, lengths 2.5-3.5 times widths, narrowly lanceolate to lanceolate, laterally compressed, usually not glaucous;

florets (1)2-5;

rachilla internodes usually shorter than 1 mm, smooth, muriculate, or scabrous, usually puberulent, infrequently hispidulous or glabrous.

Glumes

lanceolate, distinctly keeled;

lower glumes 3-veined, distinctly shorter than the lowest lemmas;

calluses glabrous or diffusely webbed, hairs less than 1/4 the lemma length;

lemmas (3)4-7 mm, lanceolate to broadly lanceolate, distinctly keeled, membranous to thinly membranous, smooth or sparsely to densely scabrous, glabrous or the keels and/or marginal veins puberulent proximally, lateral veins obscure to prominent, margins glabrous, apices acute;

palea keels scabrous, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm), aborted late in development, or 2-3.5 mm.

subulate to narrowly lanceolate, distinctly keeled, keels smooth or sparsely scabrous, apices sharply acute to acuminate;

lower glumes 3-veined, long-tapered to a slender point, lengths 6.4-11 times widths;

upper glumes shorter than or subequal to the lowest lemmas;

calluses webbed, hairs sparse, often short;

lemmas 2.4-4 mm, proximal lemma widths less than 1/5 times lengths, narrowly lanceolate to lanceolate, distinctly keeled, keels and marginal veins short-villous, lateral veins glabrous, obscure, intercostal regions smooth or muriculate, glabrous, margins glabrous, apices acute, usually partially bronze-colored;

palea keels scabrous, intercostal regions glabrous;

anthers 0.8-1.9 mm.

2n

= 28, 28+11, 56, 56+11, 59, ca. 70.

= 28, 35, 42, 48, 50, 56.

Poa cusickii

Poa nemoralis

Distribution
from FNA
CA; CO; ID; MT; ND; NV; OR; UT; WA; WY; AB; BC; MB; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa cusickii grows in rich meadows in sagebrush scrub to rocky alpine slopes, from the southwestern Yukon Territory to Manitoba and North Dakota, south to central California and eastern Colorado. It is gynodioecious or dioecious.

Sexually reproducing plants of Poa cusickii subspp. cusickii and pallida grow in different geographic areas, but pistillate plants of these two subspecies have overlapping ranges. Only pistillate plants are known in Poa cusickii subspp. epilis and purpurascens. All the alpine plants studied were pistillate.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Introduced from northern Eurasia, Poa nemoralis is established primarily at low elevations in deciduous and mixed conifer/deciduous forests. It is now common in southeastern Canada and the northeastern United States, and is spreading in the west. It can be distinguished from P. glauca (p. 576) and P. interior (see next) by its consistently short ligules, high top culm node, relatively long flag leaf blades, and narrow glumes and lemmas. It is usually hexaploid.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Panicle branches smooth or slightly scabrous, or the basal blades more than 1.5 mm wide and flat or folded; cauline blades more than 1.5 mm wide, often flat; some basal branching extravaginal; lemmas and calluses sometimes sparsely puberulent.
→ 2
2. Lemmas usually glabrous, rarely plants from the Rocky Mountains with puberulent keels and marginal veins; calluses glabrous; panicles erect, usually contracted; branches smooth to slightly scabrous
subsp. epilis
2. Lemmas rarely completely glabrous, at least some florets v/ith sparsely puberulent keels, the marginal veins glabrous or puberulent; calluses frequendy with a sparse, short web; panicles somewhat lax and loosely contracted; branches smooth or sparsely to moderately scabrous
subsp. purpurascens
1. Panicle branches moderately to strongly scabrous; basal and cauline blades usually less than 1.5 mm wide, involute, rarely flat or folded; basal branching intravaginal; lemmas and calluses glabrous.
→ 3
3. Panicle branches longer than 1.7 cm in at least some panicles; panicles open or contracted
subsp. cusickii
3. Panicle branches up to 1.7 cm long, stout; panicles contracted
subsp. pallida
Source FNA vol. 24, p. 559. FNA vol. 24, p. 574.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Epiles Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Stenopoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. cusickii subsp. cusickii, P. cusickii subsp. epilis, P. cusickii subsp. pallida, P. cusickii subsp. purpurascens
Name authority Vasey L.
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