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Cusick's bluegrass

Fendler's bluegrass, long-tongue mutton grass, mutton grass, Vasey's mutton grass

Habit Plants perennial; usually densely tufted, rarely moderately densely tufted, usually neither rhizomatous nor stoloniferous, infrequently short-rhizomatous or stoloniferous, rarely with distinct rhizomes. Plants perennial; densely to loosely tufted, rhizomatous, often weakly so, rhizomes usually short and inconspicuous.
Culms

10-60(70) cm tall, 0.5-1.8 mm thick, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-2 exserted.

15-70 cm, sometimes stout, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-1 exserted.

Sheaths

closed for 1/4-3/4 their length, terete, smooth or scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths 1.6-10 times blade lengths;

collars smooth or scabrous, glabrous;

ligules of cauline leaves 1-3(6) mm, smooth or scabrous, truncate to acute, ligules of the innovation leaves 0.2-0.5(2.5) mm, scabrous, usually truncate;

innovation blades sometimes distinctly different from the cauline blades, 0.5-2 mm wide, involute, moderately thick, moderately firm, adaxial surfaces usually densely scabrous or hispidulous to softly puberulent, infrequently nearly smooth and glabrous;

cauline blades subequal or the midcauline blades longest or the blades gradually reduced in length distally, 0.5-3 mm wide, flat, folded, or involute, usually thin, usually withering, abaxial surfaces smooth or scabrous, apices narrowly to broadly prow-shaped, flag leaf blades 0.5-5(6) cm.

closed for about 1/3 their length, terete, smooth or scabrous, glabrous or occasionally retrorsely pubescent, bases of basal sheaths glabrous, distal sheath lengths usually (5)9+ times blade lengths;

collars smooth or scabrous, glabrous or hispidulous;

ligules 0.2-18 mm, smooth or scabrous, decurrent or not, apices truncate to acuminate, ciliolate or glabrous;

innovation blades usually moderately to densely scabrous or hispidulous on and between the veins, infrequently nearly smooth and glabrous;

cauline blades strongly reduced in length distally, (0.5)1-3(4) mm wide, usually involute, moderately thick and firm, infrequently moderately thin, abaxial surfaces usually smooth, infrequently scabrous, apices narrowly prow-shaped, steeply reduced in length distally along the culm, flag leaf blades often absent or very reduced, sometimes to 1(3) cm.

Basal branching

intravaginal or intra- and extravaginal.

mainly intravaginal, usually some extravaginal.

Panicles

2-10(12) cm, usually erect, contracted or loosely contracted, narrowly lanceoloid to ovoid, congested or moderately congested, with 10-100 spikelets and 1-3(5) branches per node;

branches 0.5-4(5) cm, erect or steeply ascending, fairly straight, slender to stout, terete to angled, smooth or scabrous, with 1-15 spikelets.

2-12(30) cm, erect, contracted, narrowly lanceoloid to ovoid, congested, frequently with 100+ spikelets;

nodes with 1-2 branches;

branches 1-8 cm, erect, terete to weakly angled, smooth or scabrous, with 3-15(25) spikelets.

Spikelets

(3)4-10 mm, lengths to 3 times widths, broadly lanceolate to narrowly ovate, laterally compressed, not sexually dimorphic;

florets 2-6;

rachilla internodes 0.5-1.2 mm, smooth or scabrous.

(3)4-8(12) mm, lengths to 3 times widths, broadly lanceolate to ovate, laterally compressed, not sexually dimorphic;

florets 2-7(13);

rachilla internodes 0.8-1.3 mm, smooth, glabrous or hairy, hairs to 0.3 mm.

Glumes

lanceolate, distinctly keeled;

lower glumes 3-veined, distinctly shorter than the lowest lemmas;

calluses glabrous or diffusely webbed, hairs less than 1/4 the lemma length;

lemmas (3)4-7 mm, lanceolate to broadly lanceolate, distinctly keeled, membranous to thinly membranous, smooth or sparsely to densely scabrous, glabrous or the keels and/or marginal veins puberulent proximally, lateral veins obscure to prominent, margins glabrous, apices acute;

palea keels scabrous, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm), aborted late in development, or 2-3.5 mm.

lanceolate, distinctly keeled;

lower glumes 1-3-veined, distinctly shorter than the lowest lemmas;

calluses glabrous;

lemmas 3-6 mm, lanceolate, distinctly keeled, keels, marginal veins, and lateral veins glabrous or short- to long-villous or softly puberulent, lateral veins moderately prominent, intercostal regions softly puberulent or glabrous, smooth or sparsely scabrous, margins glabrous, apices acute;

palea keels scabrous, sometimes softly puberulent or long-villous at midlength, hairs to 0.4+ mm;

anthers vestigial (0.1-0.2 mm) or 2-3 mm.

2n

= 28, 28+11, 56, 56+11, 59, ca. 70.

= 28+11, 56, 56-58, 58-64.

Poa cusickii

Poa fendleriana

Distribution
from FNA
CA; CO; ID; MT; ND; NV; OR; UT; WA; WY; AB; BC; MB; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; BC; MB
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa cusickii grows in rich meadows in sagebrush scrub to rocky alpine slopes, from the southwestern Yukon Territory to Manitoba and North Dakota, south to central California and eastern Colorado. It is gynodioecious or dioecious.

Sexually reproducing plants of Poa cusickii subspp. cusickii and pallida grow in different geographic areas, but pistillate plants of these two subspecies have overlapping ranges. Only pistillate plants are known in Poa cusickii subspp. epilis and purpurascens. All the alpine plants studied were pistillate.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa fendleriana grows on rocky to rich slopes in sagebrush-scrub, interior chaparral, and southern (rarely northern) high plains grasslands to forests, and from desert hills to low alpine habitats. Its range extends from British Columbia to Manitoba and south to Mexico. It is one of the best spring fodder grasses in the eastern Great Basin, Colorado plateaus, and southern Rocky Mountains. It is dioecious. Each of the subspecies has regions of sexual reproduction in which staminate plants are common within populations, and extensive regions where only apomictic, pistillate plants are found. The sexual populations set little seed; the apomictic populations are highly fecund.

Poa fendleriana hybridizes with Poa cusickii subsp. pallida (p. 560). The hybrids are called P. xnematophylla (p. 562).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Panicle branches smooth or slightly scabrous, or the basal blades more than 1.5 mm wide and flat or folded; cauline blades more than 1.5 mm wide, often flat; some basal branching extravaginal; lemmas and calluses sometimes sparsely puberulent.
→ 2
2. Lemmas usually glabrous, rarely plants from the Rocky Mountains with puberulent keels and marginal veins; calluses glabrous; panicles erect, usually contracted; branches smooth to slightly scabrous
subsp. epilis
2. Lemmas rarely completely glabrous, at least some florets v/ith sparsely puberulent keels, the marginal veins glabrous or puberulent; calluses frequendy with a sparse, short web; panicles somewhat lax and loosely contracted; branches smooth or sparsely to moderately scabrous
subsp. purpurascens
1. Panicle branches moderately to strongly scabrous; basal and cauline blades usually less than 1.5 mm wide, involute, rarely flat or folded; basal branching intravaginal; lemmas and calluses glabrous.
→ 3
3. Panicle branches longer than 1.7 cm in at least some panicles; panicles open or contracted
subsp. cusickii
3. Panicle branches up to 1.7 cm long, stout; panicles contracted
subsp. pallida
1. Lemma keels and marginal veins glabrous or almost so
subsp. albescens
1. Lemma keels and marginal veins conspicuously hairy.
→ 2
2. Ligules of the middle cauline leaves 0.2-1.2 (1.5) mm long, not decurrent, usually scabrous, apices truncate to rounded, upper margins ciliolate or scabrous
subsp. fendleriana
2. Ligules of the middle cauline leaves (1.5)1.8-18 mm long, decurrent, usually smooth to sparsely scabrous, apices obtuse to acuminate, upper margins usually smooth, glabrous
subsp. longiligula
Source FNA vol. 24, p. 559. FNA vol. 24, p. 556.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Epiles Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. cusickii subsp. cusickii, P. cusickii subsp. epilis, P. cusickii subsp. pallida, P. cusickii subsp. purpurascens
P. fendleriana subsp. albescens, P. fendleriana subsp. fendleriana, P. fendleriana subsp. longiligula
Name authority Vasey (Steud.) Vasey
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