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bulbous blue grass

Hooker's bluegrass, Wheeler's blue grass

Habit Plants perennial; densely tufted, not rhizomatous, not stoloniferous. Plants perennial; densely to loosely tufted or with solitary shoots, shortly rhizomatous.
Culms

15-60 cm, erect or spreading, bases bulbous.

35-80 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 1-2 exserted.

Sheaths

closed for about 1/4 their length, terete, lowest sheaths with swollen bases;

ligules 1-3 mm, smooth or scabrous, apices obtuse to acute;

blades 1-2.5 mm wide, flat, thin, lax, soon withering.

closed for 1/3-1/4 their length, terete to slightly compressed, at least some proximal sheaths densely retrorsely scabrous, hispidulous, or softly puberulent for the upper 1/4 of their length, bases of basal sheaths glabrous, distal sheath lengths (1.4)1.7-4.6(6.2) times blade lengths;

collars of proximal leaves glabrous or with hairs the same length as those of their sheaths;

ligules 0.5-2 mm, smooth or scabrous, sometimes puberulent, truncate, those of the lower culm and innovation leaves 0.5-1.5 mm, abaxial surfaces scabrous to softly puberulent, truncate;

innovation blades folded or involute, infrequently flat, moderately thick, soft, adaxial surfaces usually densely scabrous to hispidulous;

cauline blades 2-3.5 mm wide, flat or folded, smooth or sparsely scabrous, glabrous or hispidulous, apices narrowly to broadly prow-shaped, blades gradually reduced distally or the middle blades longest, flag leaf blades 1-10 cm long.

Basal branching

intravaginal.

mainly extra-vaginal.

Panicles

3-12 cm, ovoid;

nodes with 2-5 branches;

branches ascending to spreading, terete, usually smooth or sparsely scabrous, infrequently moderately scabrous.

5-12(18) cm, erect or nodding, ovoid to pyramidal, loosely contracted to open, with 20-70 spikelets, proximal internodes usually shorter than 3.5 cm;

nodes with 2-5 branches;

branches (1)1.7-6.5 cm, ascending to spreading or reflexed, lax, terete or weakly angled, sparsely to moderately scabrous, with 2-8(12) spikelets.

Spikelets

3-5 mm, laterally compressed, usually bulbiferous;

florets 3-7, the basal floret, and sometimes additional florets, normal;

rachilla internodes smooth, glabrous.

5.5-10 mm, lengths to 3.5 times widths, laterally compressed, not sexually dimorphic;

florets 2-7;

rachilla internodes smooth or scabrous, glabrous or sparsely to densely hispidulous.

Glumes

keeled, keels scabrous;

lower glumes 3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses webbed or glabrous;

lemmas 3-4 mm, lanceolate, keeled, glabrous or the keels and marginal veins short- to long-villous, intercostal regions glabrous or softly puberulent, apices acute;

paleas scabrous, keels often softly puberulent at midlength;

anthers 1.2-1.5 mm and functional, sometimes aborted late in development, sometimes not developed.

1/4 - 2/3(3/4) as long as the adjacent lemmas, lanceolate, distinctly keeled;

lower glumes 1-3-veined, 1/4-1/2 as long as the adjacent lemmas;

calluses glabrous;

lemmas 3-6 mm, lanceolate, distinctly keeled, keels and marginal veins glabrous or softly puberulent to short-villous, intercostal regions glabrous or hispidulous, infrequently puberulent, smooth or finely scabrous, lateral veins obscure to moderately prominent, margins glabrous, apices acute;

palea keels scabrous, intercostal regions glabrous;

anthers usually vestigial (0.1-0.2 mm) or aborted late in development and up to 2 mm, rarely normal.

2n

= 14, 21, 28, 39, 42, 45.

= 56, 61, 62, 63, 64, 66, 61, 70, ca. 74, 75, 79, 80, 81, 87, 89, 90, 91.

Poa bulbosa

Poa wheeleri

Distribution
from FNA
AR; AZ; CA; CO; CT; DE; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; MB; SK
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[BONAP county map]
Discussion

Poa bulbosa is a European species that is now established in the Flora region. In southern Europe and the Middle East, it is considered an important early spring forage.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa wheeleri is common at mid- to high elevations, generally on the east side of the coastal mountains from British Columbia to California, and from Manitoba to New Mexico. It generally grows in submesic coniferous forests to subalpine habitats. Most plants have densely retrorsely pubescent or scabrous sheaths, involute innovation blades that are pubescent adaxially, and pistillate florets.

Poa wheeleri, a high polyploid apomictic species, probably arose from hybridization between P. cusickii (p. 559) and another member of the Poa nervosa complex. It resembles P. rhizomata (see next) and P. chambersii (p. 548) more than P. nervosa sensu stricto (see previous). It differs from P. chambersii in having at least some proximal sheaths that are densely retrorsely scabrous or pubescent (sometimes obscurely so), and folded or involute innovation blades that are scabrous to hispidulous on the adaxial surfaces. For a comparison with P. nervosa, see p. 545. Natural hybrids have been found between P. wheeleri and P. pratensis (p. 522).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets not bulbiferous
subsp. bulbosa
1. All or some spikelets bulbiferous
subsp. vivipara
Source FNA vol. 24, p. 516. FNA vol. 24, p. 546.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Arenariae Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. bulbosa subsp. bulbosa, P. bulbosa subsp. vivipara
Synonyms P. nervosa var. wheeleri, P. curta
Name authority L. Vasey
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