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bulbous blue grass

Douglas' bluegrass, sand dune blue grass

Habit Plants perennial; densely tufted, not rhizomatous, not stoloniferous. Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Basal branching mainly intravaginal, some extravaginal.
Culms

15-60 cm, erect or spreading, bases bulbous.

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

Sheaths

closed for about 1/4 their length, terete, lowest sheaths with swollen bases;

ligules 1-3 mm, smooth or scabrous, apices obtuse to acute;

blades 1-2.5 mm wide, flat, thin, lax, soon withering.

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

Basal branching

intravaginal.

Panicles

3-12 cm, ovoid;

nodes with 2-5 branches;

branches ascending to spreading, terete, usually smooth or sparsely scabrous, infrequently moderately scabrous.

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

Spikelets

3-5 mm, laterally compressed, usually bulbiferous;

florets 3-7, the basal floret, and sometimes additional florets, normal;

rachilla internodes smooth, glabrous.

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

Glumes

keeled, keels scabrous;

lower glumes 3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses webbed or glabrous;

lemmas 3-4 mm, lanceolate, keeled, glabrous or the keels and marginal veins short- to long-villous, intercostal regions glabrous or softly puberulent, apices acute;

paleas scabrous, keels often softly puberulent at midlength;

anthers 1.2-1.5 mm and functional, sometimes aborted late in development, sometimes not developed.

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

2n

= 14, 21, 28, 39, 42, 45.

= 28.

Poa bulbosa

Poa douglasii

Distribution
from FNA
AR; AZ; CA; CO; CT; DE; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY
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from FNA
CA
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Discussion

Poa bulbosa is a European species that is now established in the Flora region. In southern Europe and the Middle East, it is considered an important early spring forage.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets not bulbiferous
subsp. bulbosa
1. All or some spikelets bulbiferous
subsp. vivipara
Source FNA vol. 24, p. 516. FNA vol. 24, p. 551.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Arenariae Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. bulbosa subsp. bulbosa, P. bulbosa subsp. vivipara
Name authority L. Nees
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