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plains bluegrass, prairie speargrass

Hooker's bluegrass, Wheeler's blue grass

Habit Plants perennial; glaucous or not; densely to loosely tufted or the culms solitary, rhizomatous. Plants perennial; densely to loosely tufted or with solitary shoots, shortly rhizomatous.
Culms

15-80 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-1 exserted.

35-80 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 1-2 exserted.

Sheaths

closed for 1/10 – 1/5(1/4) their length, terete, smooth or sparsely scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths (1.2)1.5-9(20) times blade lengths;

ligules (1)1.5-4(5) mm, smooth or sparsely to moderately scabrous, apices obtuse to acute;

blades strongly to gradually reduced in length distally, 1.5-5 mm wide, flat and moderately thin to folded and moderately thick and firm, abaxial surfaces smooth, adaxial surfaces smooth or sparsely to moderately scabrous, primarily over the veins, apices narrowly prow-shaped, flag leaf blades (0.4)1-7(10) cm.

closed for 1/3-1/4 their length, terete to slightly compressed, at least some proximal sheaths densely retrorsely scabrous, hispidulous, or softly puberulent for the upper 1/4 of their length, bases of basal sheaths glabrous, distal sheath lengths (1.4)1.7-4.6(6.2) times blade lengths;

collars of proximal leaves glabrous or with hairs the same length as those of their sheaths;

ligules 0.5-2 mm, smooth or scabrous, sometimes puberulent, truncate, those of the lower culm and innovation leaves 0.5-1.5 mm, abaxial surfaces scabrous to softly puberulent, truncate;

innovation blades folded or involute, infrequently flat, moderately thick, soft, adaxial surfaces usually densely scabrous to hispidulous;

cauline blades 2-3.5 mm wide, flat or folded, smooth or sparsely scabrous, glabrous or hispidulous, apices narrowly to broadly prow-shaped, blades gradually reduced distally or the middle blades longest, flag leaf blades 1-10 cm long.

Basal branching

intra- and extravaginal.

mainly extra-vaginal.

Panicles

(2.5)4-12(18) cm, erect, usually narrowly lanceoloid, contracted, sometimes interrupted, infrequently loosely contracted, usually congested, with 25-100 spikelets;

nodes with 1-5 branches;

branches 1-9 cm, erect to infrequently ascending, rarely spreading, terete to weakly angled, smooth or the angles sparsely to moderately scabrous, with 3-24 spikelets.

5-12(18) cm, erect or nodding, ovoid to pyramidal, loosely contracted to open, with 20-70 spikelets, proximal internodes usually shorter than 3.5 cm;

nodes with 2-5 branches;

branches (1)1.7-6.5 cm, ascending to spreading or reflexed, lax, terete or weakly angled, sparsely to moderately scabrous, with 2-8(12) spikelets.

Spikelets

3.2-7 mm, lengths to 3.5(3.8) times widths, laterally compressed;

florets 2-7;

rachilla internodes smooth, sometimes sparsely puberulent.

5.5-10 mm, lengths to 3.5 times widths, laterally compressed, not sexually dimorphic;

florets 2-7;

rachilla internodes smooth or scabrous, glabrous or sparsely to densely hispidulous.

Glumes

lanceolate, distinctly keeled, smooth or sparsely scabrous;

lower glumes 3-veined;

calluses usually glabrous, infrequently webbed, hairs to 1/4 the lemma length;

lemmas 2.5-4.5 mm, lanceolate to narrowly lanceolate, distinctly to weakly keeled, keels and marginal veins short- to long-villous, lateral veins moderately prominent, glabrous or puberulent, intercostal regions usually glabrous, infrequently hairy, hairs to 0.3 mm, margins glabrous, apices acute or blunt;

palea keels scabrous, glabrous or short-villous at midlength, intercostal regions usually glabrous, sometimes puberulent to short-villous;

anthers 1.3-2.2 mm.

1/4 - 2/3(3/4) as long as the adjacent lemmas, lanceolate, distinctly keeled;

lower glumes 1-3-veined, 1/4-1/2 as long as the adjacent lemmas;

calluses glabrous;

lemmas 3-6 mm, lanceolate, distinctly keeled, keels and marginal veins glabrous or softly puberulent to short-villous, intercostal regions glabrous or hispidulous, infrequently puberulent, smooth or finely scabrous, lateral veins obscure to moderately prominent, margins glabrous, apices acute;

palea keels scabrous, intercostal regions glabrous;

anthers usually vestigial (0.1-0.2 mm) or aborted late in development and up to 2 mm, rarely normal.

2n

= 56, 56+1, 56-58, 63, 64, 70, 76, 84, ca. 90, 95+-5, 100, 103.

= 56, 61, 62, 63, 64, 66, 61, 70, ca. 74, 75, 79, 80, 81, 87, 89, 90, 91.

Poa arida

Poa wheeleri

Distribution
from FNA
CO; IA; IL; IN; KS; MI; MN; MT; ND; NE; NJ; NM; OH; OK; SD; TX; WY; AB; MB; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; MB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa arida grows mainly on the eastern slope of the Rocky Mountains and in the northern Great Plains, primarily in riparian habitats of varying salinity or alkalinity. It is spreading eastward along heavily salted highway corridors. Reports of its occurrence west of the Continental Divide and in southwestern Texas are mostly attributable to misidentifications of P. arctica subsp. aperta (p. 530). P. arctica subsp. grayana (p. 532), and rhizomatous specimens of P. fendleriana (p. 556).

Poa arida may reflect past hybridization between P. secunda (p. 586) and a species of Poa sect. Poa. Poa glaucifolia Scribn. & T.A. Williams refers to specimens of the northern Great Plains that have a more lax growth form with broader leaves and occasionally somewhat open panicles, florets with a small web, and sometimes lacking hairs between the keel and marginal veins of the lemma. Plants with these characteristics have chromosome counts of 2ra = 56 and 70, whereas P. arida sensu stricto usually has 2n = 63, 64, or greater than 70. It is suspected that some of the variability reflects introgression from P. secunda.

Poa arida is a named intersectional hybrid

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa wheeleri is common at mid- to high elevations, generally on the east side of the coastal mountains from British Columbia to California, and from Manitoba to New Mexico. It generally grows in submesic coniferous forests to subalpine habitats. Most plants have densely retrorsely pubescent or scabrous sheaths, involute innovation blades that are pubescent adaxially, and pistillate florets.

Poa wheeleri, a high polyploid apomictic species, probably arose from hybridization between P. cusickii (p. 559) and another member of the Poa nervosa complex. It resembles P. rhizomata (see next) and P. chambersii (p. 548) more than P. nervosa sensu stricto (see previous). It differs from P. chambersii in having at least some proximal sheaths that are densely retrorsely scabrous or pubescent (sometimes obscurely so), and folded or involute innovation blades that are scabrous to hispidulous on the adaxial surfaces. For a comparison with P. nervosa, see p. 545. Natural hybrids have been found between P. wheeleri and P. pratensis (p. 522).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 599. FNA vol. 24, p. 546.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. glaucifolia P. nervosa var. wheeleri, P. curta
Name authority Vasey Vasey
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