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plains bluegrass, prairie speargrass

ocean bluff blue grass, San Francisco bluegrass, sea-bluff bluegrass

Habit Plants perennial; glaucous or not; densely to loosely tufted or the culms solitary, rhizomatous. Plants perennial; frequently glaucous; densely tufted, not stoloniferous, not rhizomatous.
Culms

15-80 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-1 exserted.

5-40 cm, erect or ascending, frequently decumbent, terete, with 0-2 exserted nodes.

Sheaths

closed for 1/10 – 1/5(1/4) their length, terete, smooth or sparsely scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths (1.2)1.5-9(20) times blade lengths;

ligules (1)1.5-4(5) mm, smooth or sparsely to moderately scabrous, apices obtuse to acute;

blades strongly to gradually reduced in length distally, 1.5-5 mm wide, flat and moderately thin to folded and moderately thick and firm, abaxial surfaces smooth, adaxial surfaces smooth or sparsely to moderately scabrous, primarily over the veins, apices narrowly prow-shaped, flag leaf blades (0.4)1-7(10) cm.

closed for 1/10(1/5) their length, terete, smooth, glabrous, bases of basal sheaths glabrous, distal sheath lengths 1-4 times widths;

ligules 2-6 mm, smooth or sparsely scabrous, obtuse to acute;

innovation blades usually 1-1.5 mm wide, thin, soon withering, and distinctly narrower than the cauline blades, infrequently wider, flat, and a bit fleshy as in the cauline blades, or involute;

cauline blades gradually reduced in length distally, 2-5 mm wide, flat or folded, soft, thin and soon withering or moderately thick and somewhat fleshy and retaining their form, smooth, apices narrowly to broadly prow-shaped.

Basal branching

intra- and extravaginal.

all or mainly intravaginal.

Panicles

(2.5)4-12(18) cm, erect, usually narrowly lanceoloid, contracted, sometimes interrupted, infrequently loosely contracted, usually congested, with 25-100 spikelets;

nodes with 1-5 branches;

branches 1-9 cm, erect to infrequently ascending, rarely spreading, terete to weakly angled, smooth or the angles sparsely to moderately scabrous, with 3-24 spikelets.

3-7 cm, erect, nearly cylindrical, contracted, congested, with (20)30-80(120) spikelets;

nodes with 3-7 branches;

branches 0.5-1.5(4.5) cm, erect, usually angled, infrequently terete or sulcate, angles usually moderately to densely scabrous, infrequently smooth with dense papillae and weak-angled, with 2-10 spikelets in the proximal 2/3;

pedicels shorter than the spikelets.

Spikelets

3.2-7 mm, lengths to 3.5(3.8) times widths, laterally compressed;

florets 2-7;

rachilla internodes smooth, sometimes sparsely puberulent.

4.5-7 mm, lengths to 3.5 times widths, lanceolate to narrowly ovate, drab;

florets 3-5;

rachilla internodes usually shorter than 1 mm, smooth.

Glumes

lanceolate, distinctly keeled, smooth or sparsely scabrous;

lower glumes 3-veined;

calluses usually glabrous, infrequently webbed, hairs to 1/4 the lemma length;

lemmas 2.5-4.5 mm, lanceolate to narrowly lanceolate, distinctly to weakly keeled, keels and marginal veins short- to long-villous, lateral veins moderately prominent, glabrous or puberulent, intercostal regions usually glabrous, infrequently hairy, hairs to 0.3 mm, margins glabrous, apices acute or blunt;

palea keels scabrous, glabrous or short-villous at midlength, intercostal regions usually glabrous, sometimes puberulent to short-villous;

anthers 1.3-2.2 mm.

lanceolate, slightly unequal, distinctly keeled, keels papillate or scabrous;

lower glumes 3-veined;

calluses glabrous or with a crown of hairs, hairs 0.1-0.2 mm;

lemmas 3-4.5 mm, lanceolate, distinctly keeled, glabrous or the keels and marginal veins short-villous to midlength, intercostal regions sparsely puberulent near the base, margins glabrous, apices acute;

palea keels scabrous, sometimes softly puberulent at midlength;

anthers 1.5-3 mm.

2n

= 56, 56+1, 56-58, 63, 64, 70, 76, 84, ca. 90, 95+-5, 100, 103.

= 42, 84.

Poa arida

Poa unilateralis

Distribution
from FNA
CO; IA; IL; IN; KS; MI; MN; MT; ND; NE; NJ; NM; OH; OK; SD; TX; WY; AB; MB; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa arida grows mainly on the eastern slope of the Rocky Mountains and in the northern Great Plains, primarily in riparian habitats of varying salinity or alkalinity. It is spreading eastward along heavily salted highway corridors. Reports of its occurrence west of the Continental Divide and in southwestern Texas are mostly attributable to misidentifications of P. arctica subsp. aperta (p. 530). P. arctica subsp. grayana (p. 532), and rhizomatous specimens of P. fendleriana (p. 556).

Poa arida may reflect past hybridization between P. secunda (p. 586) and a species of Poa sect. Poa. Poa glaucifolia Scribn. & T.A. Williams refers to specimens of the northern Great Plains that have a more lax growth form with broader leaves and occasionally somewhat open panicles, florets with a small web, and sometimes lacking hairs between the keel and marginal veins of the lemma. Plants with these characteristics have chromosome counts of 2ra = 56 and 70, whereas P. arida sensu stricto usually has 2n = 63, 64, or greater than 70. It is suspected that some of the variability reflects introgression from P. secunda.

Poa arida is a named intersectional hybrid

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa unilateralis grows on grassy bluffs and cliffs near the Pacific coast of North America, from Washington to California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lemmas villous on the keels and marginal veins for more than 1/3 the length of the lemmas; blades usually involute, the cauline and innovation blades similar
subsp. pachypholis
1. Lemmas glabrous or the keels and marginal veins villous or puberulent for less than 1/5 the length of the lemmas; blades flat or folded, the cauline blades sometimes wider and thicker than the innovation blades
subsp. unilateralis
Source FNA vol. 24, p. 599. FNA vol. 24, p. 594.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Secundae > subsect. Halophytae
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. unilateralis subsp. pachypholis, P. unilateralis subsp. unilateralis
Synonyms P. glaucifolia
Name authority Vasey Scribn.
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