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plains bluegrass, prairie speargrass

large-glume bluegrass

Habit Plants perennial; glaucous or not; densely to loosely tufted or the culms solitary, rhizomatous. Plants perennial; green or strongly purplish; rhizomatous, rhizomes sometimes poorly developed, densely to loosely tufted.
Culms

15-80 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-1 exserted.

(20)30-120 cm, usually stout, erect or decumbent, not branching above the base, terete or weakly compressed;

nodes terete, 1-2 exserted.

Sheaths

closed for 1/10 – 1/5(1/4) their length, terete, smooth or sparsely scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths (1.2)1.5-9(20) times blade lengths;

ligules (1)1.5-4(5) mm, smooth or sparsely to moderately scabrous, apices obtuse to acute;

blades strongly to gradually reduced in length distally, 1.5-5 mm wide, flat and moderately thin to folded and moderately thick and firm, abaxial surfaces smooth, adaxial surfaces smooth or sparsely to moderately scabrous, primarily over the veins, apices narrowly prow-shaped, flag leaf blades (0.4)1-7(10) cm.

closed for 1/4 - 3/5 their length, terete to compressed, usually distinctly keeled, glabrous, sparsely to moderately scabrous, bases of basal sheaths glabrous, distal sheath lengths (0.8)1-2(2.5) times blade lengths;

collars smooth, glabrous;

ligules 2-5(6) mm, usually smooth, ligules of proximal leaves sometimes scabrous, obtuse, those of the distal leaves acute;

blades 4-12(18) cm long, (2)3-7 mm wide, flat or loosely folded, glabrous, both surfaces smooth or the adaxial surfaces sparsely scabrous, apices broadly prow-shaped, cauline blades subequal, flag leaf blades 3-10 cm.

Basal branching

intra- and extravaginal.

all or mainly extravaginal.

Panicles

(2.5)4-12(18) cm, erect, usually narrowly lanceoloid, contracted, sometimes interrupted, infrequently loosely contracted, usually congested, with 25-100 spikelets;

nodes with 1-5 branches;

branches 1-9 cm, erect to infrequently ascending, rarely spreading, terete to weakly angled, smooth or the angles sparsely to moderately scabrous, with 3-24 spikelets.

(4)7-15(20) cm, lax, loosely contracted or open, moderately congested, with (15)20-100 spikelets and 2-4(5) branches per node;

branches 3-8 cm, ascending or eventually spreading, flexuous, terete or weakly angled, usually sparsely scabridulous or scabrous, infrequently moderately densely scabrous, with (3)5-15(30) spikelets moderately crowded in the distal 1/2.

Spikelets

3.2-7 mm, lengths to 3.5(3.8) times widths, laterally compressed;

florets 2-7;

rachilla internodes smooth, sometimes sparsely puberulent.

(5)6-9(12) mm, lengths to 3 times widths, laterally compressed, sometimes bulbiferous;

florets 2-5, infrequently bulb-forming;

rachilla internodes smooth or sparsely hispidulous.

Glumes

lanceolate, distinctly keeled, smooth or sparsely scabrous;

lower glumes 3-veined;

calluses usually glabrous, infrequently webbed, hairs to 1/4 the lemma length;

lemmas 2.5-4.5 mm, lanceolate to narrowly lanceolate, distinctly to weakly keeled, keels and marginal veins short- to long-villous, lateral veins moderately prominent, glabrous or puberulent, intercostal regions usually glabrous, infrequently hairy, hairs to 0.3 mm, margins glabrous, apices acute or blunt;

palea keels scabrous, glabrous or short-villous at midlength, intercostal regions usually glabrous, sometimes puberulent to short-villous;

anthers 1.3-2.2 mm.

subequal in length and width, usually nearly equaling the adjacent lemmas, lanceolate to broadly lanceolate, distinctly keeled, keels sparsely to moderately scabrous distally, lateral veins prominent;

lower glumes (4)4.5-7 mm, 3-veined: upper glumes (4)4.5-7(8) mm, 3(5)-veined;

calluses dorsally webbed, hairs 1/3 – 2/3 times the lemma length, copious;

lemmas (4)5-8 mm, lanceolate to broadly lanceolate, green or purple, distinctly keeled, keels and marginal veins long-villous, lateral veins prominent, usually softly puberulent to short-villous, intercostal regions sometimes hispidulous, infrequently softly puberulent or short-villous proximally and hispidulous in the central 1/3, sometimes merely muriculate proximally and scabrous distally, sometimes densely scabrous throughout, rarely glabrous and nearly smooth proximally and sparsely scabrous distally, margins broadly hyaline, glabrous, apices acute;

palea keels scabrous, sometimes softly puberulent at midlength, intercostal regions broad, distinctly hispidulous or softly puberulent;

anthers 1.5-2.5 mm.

2n

= 56, 56+1, 56-58, 63, 64, 70, 76, 84, ca. 90, 95+-5, 100, 103.

= 42, 43, 44, 45, ca. 46, 49, 56, ca. 58, 63, ca. 64, ca. 65, 66, 67, 68, 69, 70, ca. 71, ca. 72, ca. 73, 74, 75, 76, 77, 78, 80, ca. 82, ca. 84, ca. 87, ca. 100.

Poa arida

Poa macrocalyx

Distribution
from FNA
CO; IA; IL; IN; KS; MI; MN; MT; ND; NE; NJ; NM; OH; OK; SD; TX; WY; AB; MB; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; BC
[BONAP county map]
Discussion

Poa arida grows mainly on the eastern slope of the Rocky Mountains and in the northern Great Plains, primarily in riparian habitats of varying salinity or alkalinity. It is spreading eastward along heavily salted highway corridors. Reports of its occurrence west of the Continental Divide and in southwestern Texas are mostly attributable to misidentifications of P. arctica subsp. aperta (p. 530). P. arctica subsp. grayana (p. 532), and rhizomatous specimens of P. fendleriana (p. 556).

Poa arida may reflect past hybridization between P. secunda (p. 586) and a species of Poa sect. Poa. Poa glaucifolia Scribn. & T.A. Williams refers to specimens of the northern Great Plains that have a more lax growth form with broader leaves and occasionally somewhat open panicles, florets with a small web, and sometimes lacking hairs between the keel and marginal veins of the lemma. Plants with these characteristics have chromosome counts of 2ra = 56 and 70, whereas P. arida sensu stricto usually has 2n = 63, 64, or greater than 70. It is suspected that some of the variability reflects introgression from P. secunda.

Poa arida is a named intersectional hybrid

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa macrocalyx grows mainly in coastal areas of boreal Alaska, and from the eastern coast of Russia to northern Japan. Bulbiferous plants are occasionally found. Poa macrocalyx resembles an exaggeratedly robust P. pratensis (p. 522), with large spikelets and lemmas, proportionally longer glumes, and paleas that are generally hispidulous between the veins and palea keels. It is cytologically and morphologically complex, and is sometimes difficult to distinguish from P. arctica subsp. lanata (see below). Poa norbergii Hulten may belong to this species; alternatively, it may be a hybrid between P. glauca (p. 576) and P. arctica (p. 576) or P. macrocalyx.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 599. FNA vol. 24, p. 527.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Poa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. glaucifolia P. turneri, P. ×norbergii, P. hispidula
Name authority Vasey Trautv. & CA. Mey.
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