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plains bluegrass, prairie speargrass

Fendler's bluegrass, long-tongue mutton grass, mutton grass, Vasey's mutton grass

Habit Plants perennial; glaucous or not; densely to loosely tufted or the culms solitary, rhizomatous. Plants perennial; densely to loosely tufted, rhizomatous, often weakly so, rhizomes usually short and inconspicuous.
Culms

15-80 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-1 exserted.

15-70 cm, sometimes stout, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-1 exserted.

Sheaths

closed for 1/10 – 1/5(1/4) their length, terete, smooth or sparsely scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths (1.2)1.5-9(20) times blade lengths;

ligules (1)1.5-4(5) mm, smooth or sparsely to moderately scabrous, apices obtuse to acute;

blades strongly to gradually reduced in length distally, 1.5-5 mm wide, flat and moderately thin to folded and moderately thick and firm, abaxial surfaces smooth, adaxial surfaces smooth or sparsely to moderately scabrous, primarily over the veins, apices narrowly prow-shaped, flag leaf blades (0.4)1-7(10) cm.

closed for about 1/3 their length, terete, smooth or scabrous, glabrous or occasionally retrorsely pubescent, bases of basal sheaths glabrous, distal sheath lengths usually (5)9+ times blade lengths;

collars smooth or scabrous, glabrous or hispidulous;

ligules 0.2-18 mm, smooth or scabrous, decurrent or not, apices truncate to acuminate, ciliolate or glabrous;

innovation blades usually moderately to densely scabrous or hispidulous on and between the veins, infrequently nearly smooth and glabrous;

cauline blades strongly reduced in length distally, (0.5)1-3(4) mm wide, usually involute, moderately thick and firm, infrequently moderately thin, abaxial surfaces usually smooth, infrequently scabrous, apices narrowly prow-shaped, steeply reduced in length distally along the culm, flag leaf blades often absent or very reduced, sometimes to 1(3) cm.

Basal branching

intra- and extravaginal.

mainly intravaginal, usually some extravaginal.

Panicles

(2.5)4-12(18) cm, erect, usually narrowly lanceoloid, contracted, sometimes interrupted, infrequently loosely contracted, usually congested, with 25-100 spikelets;

nodes with 1-5 branches;

branches 1-9 cm, erect to infrequently ascending, rarely spreading, terete to weakly angled, smooth or the angles sparsely to moderately scabrous, with 3-24 spikelets.

2-12(30) cm, erect, contracted, narrowly lanceoloid to ovoid, congested, frequently with 100+ spikelets;

nodes with 1-2 branches;

branches 1-8 cm, erect, terete to weakly angled, smooth or scabrous, with 3-15(25) spikelets.

Spikelets

3.2-7 mm, lengths to 3.5(3.8) times widths, laterally compressed;

florets 2-7;

rachilla internodes smooth, sometimes sparsely puberulent.

(3)4-8(12) mm, lengths to 3 times widths, broadly lanceolate to ovate, laterally compressed, not sexually dimorphic;

florets 2-7(13);

rachilla internodes 0.8-1.3 mm, smooth, glabrous or hairy, hairs to 0.3 mm.

Glumes

lanceolate, distinctly keeled, smooth or sparsely scabrous;

lower glumes 3-veined;

calluses usually glabrous, infrequently webbed, hairs to 1/4 the lemma length;

lemmas 2.5-4.5 mm, lanceolate to narrowly lanceolate, distinctly to weakly keeled, keels and marginal veins short- to long-villous, lateral veins moderately prominent, glabrous or puberulent, intercostal regions usually glabrous, infrequently hairy, hairs to 0.3 mm, margins glabrous, apices acute or blunt;

palea keels scabrous, glabrous or short-villous at midlength, intercostal regions usually glabrous, sometimes puberulent to short-villous;

anthers 1.3-2.2 mm.

lanceolate, distinctly keeled;

lower glumes 1-3-veined, distinctly shorter than the lowest lemmas;

calluses glabrous;

lemmas 3-6 mm, lanceolate, distinctly keeled, keels, marginal veins, and lateral veins glabrous or short- to long-villous or softly puberulent, lateral veins moderately prominent, intercostal regions softly puberulent or glabrous, smooth or sparsely scabrous, margins glabrous, apices acute;

palea keels scabrous, sometimes softly puberulent or long-villous at midlength, hairs to 0.4+ mm;

anthers vestigial (0.1-0.2 mm) or 2-3 mm.

2n

= 56, 56+1, 56-58, 63, 64, 70, 76, 84, ca. 90, 95+-5, 100, 103.

= 28+11, 56, 56-58, 58-64.

Poa arida

Poa fendleriana

Distribution
from FNA
CO; IA; IL; IN; KS; MI; MN; MT; ND; NE; NJ; NM; OH; OK; SD; TX; WY; AB; MB; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; BC; MB
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa arida grows mainly on the eastern slope of the Rocky Mountains and in the northern Great Plains, primarily in riparian habitats of varying salinity or alkalinity. It is spreading eastward along heavily salted highway corridors. Reports of its occurrence west of the Continental Divide and in southwestern Texas are mostly attributable to misidentifications of P. arctica subsp. aperta (p. 530). P. arctica subsp. grayana (p. 532), and rhizomatous specimens of P. fendleriana (p. 556).

Poa arida may reflect past hybridization between P. secunda (p. 586) and a species of Poa sect. Poa. Poa glaucifolia Scribn. & T.A. Williams refers to specimens of the northern Great Plains that have a more lax growth form with broader leaves and occasionally somewhat open panicles, florets with a small web, and sometimes lacking hairs between the keel and marginal veins of the lemma. Plants with these characteristics have chromosome counts of 2ra = 56 and 70, whereas P. arida sensu stricto usually has 2n = 63, 64, or greater than 70. It is suspected that some of the variability reflects introgression from P. secunda.

Poa arida is a named intersectional hybrid

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa fendleriana grows on rocky to rich slopes in sagebrush-scrub, interior chaparral, and southern (rarely northern) high plains grasslands to forests, and from desert hills to low alpine habitats. Its range extends from British Columbia to Manitoba and south to Mexico. It is one of the best spring fodder grasses in the eastern Great Basin, Colorado plateaus, and southern Rocky Mountains. It is dioecious. Each of the subspecies has regions of sexual reproduction in which staminate plants are common within populations, and extensive regions where only apomictic, pistillate plants are found. The sexual populations set little seed; the apomictic populations are highly fecund.

Poa fendleriana hybridizes with Poa cusickii subsp. pallida (p. 560). The hybrids are called P. xnematophylla (p. 562).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lemma keels and marginal veins glabrous or almost so
subsp. albescens
1. Lemma keels and marginal veins conspicuously hairy.
→ 2
2. Ligules of the middle cauline leaves 0.2-1.2 (1.5) mm long, not decurrent, usually scabrous, apices truncate to rounded, upper margins ciliolate or scabrous
subsp. fendleriana
2. Ligules of the middle cauline leaves (1.5)1.8-18 mm long, decurrent, usually smooth to sparsely scabrous, apices obtuse to acuminate, upper margins usually smooth, glabrous
subsp. longiligula
Source FNA vol. 24, p. 599. FNA vol. 24, p. 556.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. fendleriana subsp. albescens, P. fendleriana subsp. fendleriana, P. fendleriana subsp. longiligula
Synonyms P. glaucifolia
Name authority Vasey (Steud.) Vasey
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