Poa arida |
Poa chambersii |
|
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plains bluegrass, prairie speargrass |
Chambers' bluegrass |
|
Habit | Plants perennial; glaucous or not; densely to loosely tufted or the culms solitary, rhizomatous. | Plants perennial; loosely tufted or with solitary shoots, short-rhizomatous. |
Culms | 15-80 cm, erect or the bases decumbent, terete or weakly compressed; nodes terete, 0-1 exserted. |
10-50 cm, erect or the bases decumbent, terete or weakly compressed; nodes terete, 0-1 exserted. |
Sheaths | closed for 1/10 – 1/5(1/4) their length, terete, smooth or sparsely scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths (1.2)1.5-9(20) times blade lengths; ligules (1)1.5-4(5) mm, smooth or sparsely to moderately scabrous, apices obtuse to acute; blades strongly to gradually reduced in length distally, 1.5-5 mm wide, flat and moderately thin to folded and moderately thick and firm, abaxial surfaces smooth, adaxial surfaces smooth or sparsely to moderately scabrous, primarily over the veins, apices narrowly prow-shaped, flag leaf blades (0.4)1-7(10) cm. |
closed for 1/3 – 7/8 their length, terete to slightly compressed, smooth, glabrous, bases of basal sheaths glabrous, distal sheath lengths (1.15)1.5-4.6(6.6) times blade lengths; collars smooth, glabrous; ligules 0.5-2(2.5) mm, smooth, truncate to obtuse; innovation blades similar to the cauline blades; cauline blades gradually reduced in length distally, 2-5 mm wide, flat or folded, smooth or the adaxial surfaces sparsely scabrous, primarily over the veins, apices broadly prow-shaped, flag leaf blades 0.7-6 cm. |
Basal branching | intra- and extravaginal. |
all or mainly extravaginal. |
Panicles | (2.5)4-12(18) cm, erect, usually narrowly lanceoloid, contracted, sometimes interrupted, infrequently loosely contracted, usually congested, with 25-100 spikelets; nodes with 1-5 branches; branches 1-9 cm, erect to infrequently ascending, rarely spreading, terete to weakly angled, smooth or the angles sparsely to moderately scabrous, with 3-24 spikelets. |
2-9 cm, erect, lanceoloid to ovoid, tightly to loosely contracted, with 15-35 spikelets, proximal internodes shorter than 2 cm; nodes with 1-2 branches; branches 0.9-3.2 cm, erect to ascending or slightly spreading, terete, smooth or sparsely scabrous, with 1-4 spikelets. |
Spikelets | 3.2-7 mm, lengths to 3.5(3.8) times widths, laterally compressed; florets 2-7; rachilla internodes smooth, sometimes sparsely puberulent. |
6-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic; florets 2-7; rachilla internodes 0.8-1.5 mm, smooth or sparsely scabrous, glabrous. |
Glumes | lanceolate, distinctly keeled, smooth or sparsely scabrous; lower glumes 3-veined; calluses usually glabrous, infrequently webbed, hairs to 1/4 the lemma length; lemmas 2.5-4.5 mm, lanceolate to narrowly lanceolate, distinctly to weakly keeled, keels and marginal veins short- to long-villous, lateral veins moderately prominent, glabrous or puberulent, intercostal regions usually glabrous, infrequently hairy, hairs to 0.3 mm, margins glabrous, apices acute or blunt; palea keels scabrous, glabrous or short-villous at midlength, intercostal regions usually glabrous, sometimes puberulent to short-villous; anthers 1.3-2.2 mm. |
3/5- 4/5 as long as the adjacent lemmas, distinctly keeled; lower glumes 3-veined; calluses of at least some proximal florets sparsely webbed, with 1-2 mm hairs, others glabrous, rarely all glabrous; lemmas 5-7 mm, lanceolate, 5-7-veined, distinctly keeled, smooth or sparsely finely scabrous, glabrous throughout or the keels and marginal veins sparsely softly puberulent over the proximal 1/4, lateral veins moderately prominent, intercostal regions glabrous, margins glabrous, apices acute; palea keels sparsely scabrous, intercostal regions glabrous; anthers vestigial (0.1-0.2 mm), aborted late in development, or 1.8-3.7 mm. |
2n | = 56, 56+1, 56-58, 63, 64, 70, 76, 84, ca. 90, 95+-5, 100, 103. |
= unknown. |
Poa arida |
Poa chambersii |
|
Distribution |
CO; IA; IL; IN; KS; MI; MN; MT; ND; NE; NJ; NM; OH; OK; SD; TX; WY; AB; MB; ON; QC; SK
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OR |
Discussion | Poa arida grows mainly on the eastern slope of the Rocky Mountains and in the northern Great Plains, primarily in riparian habitats of varying salinity or alkalinity. It is spreading eastward along heavily salted highway corridors. Reports of its occurrence west of the Continental Divide and in southwestern Texas are mostly attributable to misidentifications of P. arctica subsp. aperta (p. 530). P. arctica subsp. grayana (p. 532), and rhizomatous specimens of P. fendleriana (p. 556). Poa arida may reflect past hybridization between P. secunda (p. 586) and a species of Poa sect. Poa. Poa glaucifolia Scribn. & T.A. Williams refers to specimens of the northern Great Plains that have a more lax growth form with broader leaves and occasionally somewhat open panicles, florets with a small web, and sometimes lacking hairs between the keel and marginal veins of the lemma. Plants with these characteristics have chromosome counts of 2ra = 56 and 70, whereas P. arida sensu stricto usually has 2n = 63, 64, or greater than 70. It is suspected that some of the variability reflects introgression from P. secunda. Poa arida is a named intersectional hybrid (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Poa chambersii is known only from upland forest openings in the Cascades of western Oregon, where it is dioecious, and from high elevations on Steens Mountain in southeastern Oregon, where it is gynodioecious. It resembles P. rhizomata (see previous), but has more closed sheaths, shorter ligules, less pubescent or glabrous lemmas, and lacks a well-developed web. It approaches P. cusickii subsp. purpurascens (p. 562), but is rhizomatous and sexually reproducing. It differs from P. wheeleri (p. 546) in having glabrous sheaths and flat or folded, glabrous innovation blades. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 24, p. 599. | FNA vol. 24, p. 548. |
Parent taxa | Poaceae > subfam. Pooideae > tribe Poeae > Poa | Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa |
Sibling taxa | ||
Synonyms | P. glaucifolia | |
Name authority | Vasey | Soreng |
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