Poa arctica |
Poa tenerrima |
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arctic bluegrass |
delicate blue grass |
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Habit | Plants perennial; usually strongly anthocyanic; rhizomatous, rhizomes usually well developed, sometimes poorly developed, shoots usually solitary. | Plants perennial; densely tufted, basal leaf tufts 2-8(13) cm, small, narrowly based, not stoloniferous, not rhizomatous. | ||||||||||||||||
Culms | 7.5-60 cm, slender to stout, terete or weakly compressed, bases usually decumbent, not branching above the bases; nodes terete, proximal nodes usually not exserted, 0-2 exserted above. |
15-50 cm tall, slender, 0.8-0.9 mm thick; nodes terete, 0-1 exserted. |
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Sheaths | closed for (1/6)1/5 – 2/5 their length, terete, glabrous, smooth or sparsely scabrous, bases of basal sheaths glabrous, distal sheath lengths 1.4-4(5.3) times blade lengths; collars smooth, glabrous; ligules (1)2-7 mm, glabrous, smooth or sparsely to infrequently moderately scabrous, apices usually rounded to obtuse or acute, rarely truncate, entire or lacerate; blades 1-6 mm wide, flat or folded, somewhat involute, smooth, glabrous, apices broadly prow-shaped, cauline blades subequal or gradually reduced distally, flag leaf blades 0.7-9 cm. |
closed for 76-74 their length, terete, scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths 2.3-7.7 times blade lengths; collars sparsely scabrous, glabrous; ligules 0.5-1.5(2.5) mm, scabrous, apices usually truncate to obtuse, sometimes acute, ligules of innovations to 0.5 mm; innovation blades filiform; cauline blades gradually reduced in length upwards, 0.4-1.5 mm wide, mostly folded, thin, soft, soon withering, scabrous, apices narrowly prow-shaped. |
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Basal branching | mainly extravaginal. |
intravaginal. |
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Panicles | (2)3.5-15 cm, ovoid to broadly pyramidal, usually open, sparse, with 10-40(60) spikelets, proximal internodes shorter than 1.5(3) cm, with (1)2-5 branches per node; branches 1.5-6 cm, spreading soon after emergence from the sheath, thin, sinuous, and flexuous to fairly stout and straight, terete, smooth or sparsely to infrequently moderately scabrous, with (1)2-5 spikelets, the spikelets not crowded. |
5-15 cm, 1.3-2.2 times the branch lengths, erect, broadly rhomboidal to pyramidal, open, purple, sparse, proximal internodes 1.5-4.2 cm; nodes with 1-2(5) branches; branches 3-8.5 cm, widely spreading, capillary, straight, terete to weakly angled, moderately to mostly densely scabrous, with 3-9 spikelets in the distal 73. |
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Spikelets | (3.5) 4.5-8 mm, lengths to 3.5 times widths, laterally compressed, sometimes bulbiferous; florets (2)3-6, infrequently bulb-forming; rachilla internodes smooth or muriculate, proximal internodes glabrous or sparsely softly puberulent to long-villous. |
5-8 mm, lengths (3.8)4-5 times widths, usually narrowly lanceolate to lanceolate, subterete to weakly laterally compressed, drab, usually strongly anthocyanic; florets 3-5; rachilla internodes 1-1.5+ mm, terete or slightly dorsally compressed, muriculate or scabrous. |
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Glumes | lanceolate to broadly lanceolate, distinctly or weakly keeled, keels usually smooth, sometimes sparsely scabrous distally, lateral veins usually moderately pronounced; lower glumes (3)3.5-5(6) mm, 3-veined; upper glumes 3.5-5.5(6.5) mm, nearly equaling to slightly exceeding the lowest lemmas, or distinctly shorter; calluses glabrous or webbed, hairs sparse and short to over 1/3 – 2/3 the lemma length; lemmas (2.7)3-6(7) mm, lanceolate to broadly lanceolate, usually strongly purple, distinctly keeled, keels, marginal veins, and lateral veins long-villous, hairs on the lateral veins sometimes shorter, lateral veins prominent, intercostal regions short-villous to softly puberulent at least near the base, glabrous elsewhere, smooth to weakly muriculate and/or usually sparsely scabrous, infrequently moderately scabrous, margins broadly hyaline, glabrous, apices acute; palea keels usually short- to long-villous for most of their length, rarely nearly glabrous and scabrous, intercostal regions broad, usually at least sparsely softly puberulent, rarely glabrous, apices scabrous; anthers 1.4-2.5 mm, sometimes aborted late in development. |
lanceolate, distinctly shorter than the adjacent lemmas, scabrous distally, keels indistinct, scabrous, obtuse to acute; lower glumes 3-veined; calluses glabrous or with a crown of hairs, hairs to 0.3 mm; lemmas 3-4.2 mm, lanceolate, weakly keeled, keels, veins, and proximal 2/3 of the intercostal regions puberulent, lateral veins obscure, margins strongly inrolled below, broadly scarious above, glabrous, apices obtuse to acute; palea keels scabrous, sometimes hairy at midlength; anthers 1.6-2.1 mm. |
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2n | = 36, 42, 56, 60, 62-68, 70, ca. 72, 74-76, 78-80, 82-84, 86, 88, 99, 106. |
= 42. |
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Poa arctica |
Poa tenerrima |
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Distribution |
AK; CO; ID; MT; NM; NV; UT; WA; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT; Greenland
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CA |
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Discussion | Poa arctica is a common circumboreal species of arctic and alpine regions, growing mainly in mesic to subhydric, acidic tundra and alpine meadows, and on rocky slopes. It extends south in the Rocky Mountains to New Mexico. The frequency of sterile anthers in plants of the high arctic suggests that P. arctica is sometimes apomictic in that region. Over most of the rest of its range, P. arctica usually develops normal anthers. This and isozyme data for populations from alpine and low arctic regions suggest sexual reproduction is common in these habitats. The most reliable way to distinguish Poa arctica from P. pratensis (p. 522), particularly subsp. alpigena, is by the wider paleas and the presence of hairs between the palea keels. Bulbiferous forms of P. arctica differ from P. stenantha var. vivipara (p. 594) in not being glaucous, and in having rhizomes and terete, smooth panicle branches. Poa xgaspensis (p. 601) also resembles P. arctica, but it has sharply keeled, more scabrous glumes and a spikelet shape that is intermediate between P. pratensis and P. alpina (p. 518). Poa arctica forms natural hybrids with both P. pratensis and P. secunda (p. 586). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Poa tenerrima is a rare species, endemic to serpentine barrens along the western base of the Sierra Nevada. It differs from P. secunda subsp. secunda (see previous) in combining consistently wide, open panicles, very scabrous branches, short-truncate ligules, and very fine foliage. A series of small, delicate, open-panicled plants from the California Coast Ranges, formerly included in P. tenerrima by Soreng (1993), differ in having smooth branches and longer ligules, and are better referred to P. secunda subsp. secunda. No intergradation is evident where the two taxa grow together. Plants from the Columbia River Gorge in Oregon, including the type of P. multnomae Piper, approach P. tenerrima, but are presumed to be derived from hybridization between P. nervosa (p. 545) and P. secunda (p. 586). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24. | FNA vol. 24, p. 588. | ||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Poa | Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Secundae > subsect. Secundae | ||||||||||||||||
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Name authority | R. Br. | Scribn. | ||||||||||||||||
Web links |