Poa arctica |
Poa sierrae |
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arctic bluegrass |
Sierra blue grass |
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Habit | Plants perennial; usually strongly anthocyanic; rhizomatous, rhizomes usually well developed, sometimes poorly developed, shoots usually solitary. | Plants perennial; loosely tufted or with solitary shoots, short-rhizomatous. | ||||||||||||||||
Culms | 7.5-60 cm, slender to stout, terete or weakly compressed, bases usually decumbent, not branching above the bases; nodes terete, proximal nodes usually not exserted, 0-2 exserted above. |
20-60 cm, slender, erect or the bases decumbent, terete or weakly compressed; nodes terete, 1-2 exserted. |
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Sheaths | closed for (1/6)1/5 – 2/5 their length, terete, glabrous, smooth or sparsely scabrous, bases of basal sheaths glabrous, distal sheath lengths 1.4-4(5.3) times blade lengths; collars smooth, glabrous; ligules (1)2-7 mm, glabrous, smooth or sparsely to infrequently moderately scabrous, apices usually rounded to obtuse or acute, rarely truncate, entire or lacerate; blades 1-6 mm wide, flat or folded, somewhat involute, smooth, glabrous, apices broadly prow-shaped, cauline blades subequal or gradually reduced distally, flag leaf blades 0.7-9 cm. |
closed from 9/10 their length to their entire length, terete, smooth or sparsely scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.18-0.8 times blade lengths; collars smooth, glabrous; ligules 3-6 mm, scabrous, acute to acuminate; innovation blades similar to the cauline blades; cauline blades gradually reduced in length distally, 1.5-2.5 mm wide, flat, thin, soft, smooth or sparsely scabrous, primarily over the veins, apices narrowly to broadly prow-shaped, flag leaf blades 8-12 cm. |
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Basal branching | mainly extravaginal. |
extravaginal, initiated as pinkish to purplish fleshy buds that persist as sets of short scales at the nodes of rhizomes and the proximal culm nodes, drying brownish and flabelliform after the shoots develop. |
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Panicles | (2)3.5-15 cm, ovoid to broadly pyramidal, usually open, sparse, with 10-40(60) spikelets, proximal internodes shorter than 1.5(3) cm, with (1)2-5 branches per node; branches 1.5-6 cm, spreading soon after emergence from the sheath, thin, sinuous, and flexuous to fairly stout and straight, terete, smooth or sparsely to infrequently moderately scabrous, with (1)2-5 spikelets, the spikelets not crowded. |
4-15 cm, erect, ovoid, sparse, with fewer than 15(20) spikelets; nodes with 1-2 branches; branches 1-4.5 cm, spreading to reflexed, slender, terete, sparsely to moderately scabrous, with 1-3 spikelets. |
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Spikelets | (3.5) 4.5-8 mm, lengths to 3.5 times widths, laterally compressed, sometimes bulbiferous; florets (2)3-6, infrequently bulb-forming; rachilla internodes smooth or muriculate, proximal internodes glabrous or sparsely softly puberulent to long-villous. |
5-9 mm, lengths to 3.5 times widths, laterally compressed, not sexually dimorphic; florets 2-6; rachilla internodes smooth, sparsely hairy, hairs to 0.3 mm. |
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Glumes | lanceolate to broadly lanceolate, distinctly or weakly keeled, keels usually smooth, sometimes sparsely scabrous distally, lateral veins usually moderately pronounced; lower glumes (3)3.5-5(6) mm, 3-veined; upper glumes 3.5-5.5(6.5) mm, nearly equaling to slightly exceeding the lowest lemmas, or distinctly shorter; calluses glabrous or webbed, hairs sparse and short to over 1/3 – 2/3 the lemma length; lemmas (2.7)3-6(7) mm, lanceolate to broadly lanceolate, usually strongly purple, distinctly keeled, keels, marginal veins, and lateral veins long-villous, hairs on the lateral veins sometimes shorter, lateral veins prominent, intercostal regions short-villous to softly puberulent at least near the base, glabrous elsewhere, smooth to weakly muriculate and/or usually sparsely scabrous, infrequently moderately scabrous, margins broadly hyaline, glabrous, apices acute; palea keels usually short- to long-villous for most of their length, rarely nearly glabrous and scabrous, intercostal regions broad, usually at least sparsely softly puberulent, rarely glabrous, apices scabrous; anthers 1.4-2.5 mm, sometimes aborted late in development. |
1/3 – 3/4(4/5) as long as the adjacent lemmas, keels sparsely scabrous; lower glumes 3-veined; calluses glabrous or webbed, hairs at least 1-2 mm; lemmas 4-7 mm, lanceolate, distinctly keeled, keels and marginal veins glabrous or short-villous, lateral veins obscure, glabrous, finely scabrous, intercostal regions glabrous or sparsely softly puberulent, margins glabrous, apices acute; palea keels scabrous, sometimes softly puberulent at midlength; anthers vestigial (0.1-0.2 mm) or 2-4 mm. |
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2n | = 36, 42, 56, 60, 62-68, 70, ca. 72, 74-76, 78-80, 82-84, 86, 88, 99, 106. |
= ca. 58. |
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Poa arctica |
Poa sierrae |
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Distribution |
AK; CO; ID; MT; NM; NV; UT; WA; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT; Greenland
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CA |
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Discussion | Poa arctica is a common circumboreal species of arctic and alpine regions, growing mainly in mesic to subhydric, acidic tundra and alpine meadows, and on rocky slopes. It extends south in the Rocky Mountains to New Mexico. The frequency of sterile anthers in plants of the high arctic suggests that P. arctica is sometimes apomictic in that region. Over most of the rest of its range, P. arctica usually develops normal anthers. This and isozyme data for populations from alpine and low arctic regions suggest sexual reproduction is common in these habitats. The most reliable way to distinguish Poa arctica from P. pratensis (p. 522), particularly subsp. alpigena, is by the wider paleas and the presence of hairs between the palea keels. Bulbiferous forms of P. arctica differ from P. stenantha var. vivipara (p. 594) in not being glaucous, and in having rhizomes and terete, smooth panicle branches. Poa xgaspensis (p. 601) also resembles P. arctica, but it has sharply keeled, more scabrous glumes and a spikelet shape that is intermediate between P. pratensis and P. alpina (p. 518). Poa arctica forms natural hybrids with both P. pratensis and P. secunda (p. 586). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Poa sierrae, a distinctive dioecious species, is a narrow endemic of mid-elevation canyon slopes on the west side of the Sierra Nevada, California. It can be distinguished from all other Poa species by the scaly, pink to purplish buds on the rhizomes, and by the entirely or almost entirely closed upper culm sheaths that are shorter than their blades. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24. | FNA vol. 24, p. 548. | ||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Poa | Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa | ||||||||||||||||
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Name authority | R. Br. | J.T. Howell | ||||||||||||||||
Web links |