Poa arctica |
Poa glauca |
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arctic bluegrass |
glaucous blue grass, timberline bluegrass, white bluegrass |
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Habit | Plants perennial; usually strongly anthocyanic; rhizomatous, rhizomes usually well developed, sometimes poorly developed, shoots usually solitary. | Plants perennial; usually glaucous; densely tufted, not stoloniferous, not rhizomatous. | ||||||||||||||||||||||||
Culms | 7.5-60 cm, slender to stout, terete or weakly compressed, bases usually decumbent, not branching above the bases; nodes terete, proximal nodes usually not exserted, 0-2 exserted above. |
5-40(80) cm, erect to spreading, straight, wiry, bases straight or slightly decumbent; nodes terete or slightly compressed, usually 0-1 exserted, top node at 1/10-1/3 the culm length. |
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Sheaths | closed for (1/6)1/5 – 2/5 their length, terete, glabrous, smooth or sparsely scabrous, bases of basal sheaths glabrous, distal sheath lengths 1.4-4(5.3) times blade lengths; collars smooth, glabrous; ligules (1)2-7 mm, glabrous, smooth or sparsely to infrequently moderately scabrous, apices usually rounded to obtuse or acute, rarely truncate, entire or lacerate; blades 1-6 mm wide, flat or folded, somewhat involute, smooth, glabrous, apices broadly prow-shaped, cauline blades subequal or gradually reduced distally, flag leaf blades 0.7-9 cm. |
closed for 1/10 – 1/5 their length, terete, bases of basal sheaths glabrous or sparsely minutely hairy, hairs 0.1-0.2 mm, distal sheath lengths 1.1-4 times blade lengths; ligules 1-4(5) mm, sparsely to densely scabrous, apices obtuse to acute, minutely ciliolate; blades 0.8-2.5 mm wide, flat or folded, thin, soft, appressed or abruptly ascending to spreading, straight, apices narrowly prow-shaped. |
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Basal branching | mainly extravaginal. |
all or mostly extravaginal. |
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Panicles | (2)3.5-15 cm, ovoid to broadly pyramidal, usually open, sparse, with 10-40(60) spikelets, proximal internodes shorter than 1.5(3) cm, with (1)2-5 branches per node; branches 1.5-6 cm, spreading soon after emergence from the sheath, thin, sinuous, and flexuous to fairly stout and straight, terete, smooth or sparsely to infrequently moderately scabrous, with (1)2-5 spikelets, the spikelets not crowded. |
1-10(20) cm, lengths 3-5 times widths at maturity, rarely racemelike with branches of irregular length, erect, narrowly lanceoloid to ovoid, contracted to somewhat open, sparse, proximal internodes shorter than 1.5(4) cm; nodes with 2-3(5) branches; branches erect, ascending or weakly spreading, fairly straight, short, stout, angled, angles moderately to densely scabrous, rarely only scabrous distally, glaucous; pedicels usually shorter than the spikelets. |
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Spikelets | (3.5) 4.5-8 mm, lengths to 3.5 times widths, laterally compressed, sometimes bulbiferous; florets (2)3-6, infrequently bulb-forming; rachilla internodes smooth or muriculate, proximal internodes glabrous or sparsely softly puberulent to long-villous. |
3-7(9) mm, lengths 2-3 times widths, laterally compressed, rarely bulbiferous, usually glaucous; florets 2-5, rarely bulb-forming; rachilla internodes to 1.2 mm, smooth, muriculate, or scabrous, glabrous or sparsely to densely hispidulous or puberulent. |
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Glumes | lanceolate to broadly lanceolate, distinctly or weakly keeled, keels usually smooth, sometimes sparsely scabrous distally, lateral veins usually moderately pronounced; lower glumes (3)3.5-5(6) mm, 3-veined; upper glumes 3.5-5.5(6.5) mm, nearly equaling to slightly exceeding the lowest lemmas, or distinctly shorter; calluses glabrous or webbed, hairs sparse and short to over 1/3 – 2/3 the lemma length; lemmas (2.7)3-6(7) mm, lanceolate to broadly lanceolate, usually strongly purple, distinctly keeled, keels, marginal veins, and lateral veins long-villous, hairs on the lateral veins sometimes shorter, lateral veins prominent, intercostal regions short-villous to softly puberulent at least near the base, glabrous elsewhere, smooth to weakly muriculate and/or usually sparsely scabrous, infrequently moderately scabrous, margins broadly hyaline, glabrous, apices acute; palea keels usually short- to long-villous for most of their length, rarely nearly glabrous and scabrous, intercostal regions broad, usually at least sparsely softly puberulent, rarely glabrous, apices scabrous; anthers 1.4-2.5 mm, sometimes aborted late in development. |
subequal, narrowly to broadly lanceolate, distinctly keeled, keels smooth or sparsely scabrous, apices acute; lower glumes 3-veined; upper glumes 2-3.8(5.2) mm, lengths usually more than 4.1 times widths, distinctly shorter to subequal to the lowest lemmas; calluses glabrous or webbed, webs from minute to more than 1/2 the lemma length; lemmas 2.5-4 mm, lanceolate to broadly lanceolate, distinctly keeled, keels and marginal veins short-villous, lateral veins obscure, usually sparsely softly puberulent to short-villous, intercostal regions smooth, sometimes weakly muriculate, glabrous or puberulent, margins glabrous, apices usually partially bronze-colored, obtuse or acute; palea keels scabrous, glabrous or softly puberulent at midlength, intercostal regions glabrous or softly puberulent; anthers (1)1.2-2.5 mm, mature sacs 0.2 mm wide, rarely aborted late in development. |
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2n | = 36, 42, 56, 60, 62-68, 70, ca. 72, 74-76, 78-80, 82-84, 86, 88, 99, 106. |
= 34, 42, 44, 47, 48, 49, 50, 56, 56, 57, 58, 60, 63, 64, 65, 70, 75, 78, ca. 100. |
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Poa arctica |
Poa glauca |
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Distribution |
AK; CO; ID; MT; NM; NV; UT; WA; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT; Greenland
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AK; AZ; CA; CO; ID; ME; MI; MN; MT; NH; NM; NV; NY; OR; PA; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Greenland
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Discussion | Poa arctica is a common circumboreal species of arctic and alpine regions, growing mainly in mesic to subhydric, acidic tundra and alpine meadows, and on rocky slopes. It extends south in the Rocky Mountains to New Mexico. The frequency of sterile anthers in plants of the high arctic suggests that P. arctica is sometimes apomictic in that region. Over most of the rest of its range, P. arctica usually develops normal anthers. This and isozyme data for populations from alpine and low arctic regions suggest sexual reproduction is common in these habitats. The most reliable way to distinguish Poa arctica from P. pratensis (p. 522), particularly subsp. alpigena, is by the wider paleas and the presence of hairs between the palea keels. Bulbiferous forms of P. arctica differ from P. stenantha var. vivipara (p. 594) in not being glaucous, and in having rhizomes and terete, smooth panicle branches. Poa xgaspensis (p. 601) also resembles P. arctica, but it has sharply keeled, more scabrous glumes and a spikelet shape that is intermediate between P. pratensis and P. alpina (p. 518). Poa arctica forms natural hybrids with both P. pratensis and P. secunda (p. 586). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Poa glauca is a common, highly variable, circumboreal, boreal forest to alpine and high arctic species. It grows from Alaska to Greenland, south to California and New Mexico in the west, and through Canada and the northeastern United States in the east. It also grows at scattered locations in Patagonia. It generally favors dry habitats and tolerates disturbance well. It can be distinguished from P. nemoralis (p. 574) and P. interior (see previous) by its longer ligules, lower top culm node, and wider glumes and lemmas. It can be difficult to distinguish from P. laxa subsp. banffiana (p. 570). Poa glauca is often confused in herbaria with P. abbreviata subsp. pattersonii (p. 582). It differs in having primarily extravaginal branching and, usually, longer anthers. It hybridizes with P. laxa, forming P. laxa x glauca (p. 572). It is also known to hybridize with P. hartzii (p. 589), and is suspected to hybridize with P. arctica (p. 529) and P. secunda (p. 586). It is highly polyploid, and presumed to be highly apomictic. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24. | FNA vol. 24, p. 576. | ||||||||||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Poa | Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Stenopoa | ||||||||||||||||||||||||
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Name authority | R. Br. | Vahl | ||||||||||||||||||||||||
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