Poa arctica
Poa diaboli
arctic bluegrass
Diablo bluegrass, Diablo Canyon blue grass
7.5-60 cm, slender to stout, terete or weakly compressed, bases usually decumbent, not branching above the bases;
nodes terete, proximal nodes usually not exserted, 0-2 exserted above.
26-50 cm tall, 0.5-0.9 mm thick, bases decumbent or nearly erect, frequently branching above the base, terete or weakly compressed;
nodes terete, 1-2 exserted.
closed for (1/6)1/5 – 2/5 their length, terete, glabrous, smooth or sparsely scabrous, bases of basal sheaths glabrous, distal sheath lengths 1.4-4(5.3) times blade lengths;
collars smooth, glabrous;
ligules (1)2-7 mm, glabrous, smooth or sparsely to infrequently moderately scabrous, apices usually rounded to obtuse or acute, rarely truncate, entire or lacerate;
blades 1-6 mm wide, flat or folded, somewhat involute, smooth, glabrous, apices broadly prow-shaped, cauline blades subequal or gradually reduced distally, flag leaf blades 0.7-9 cm.
closed for 2/5 – 7/10 their length, weakly keeled, sparsely scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.6-2.4 times blade lengths;
collars scabrous or pubescent on the margins;
ligules (1)2-3 mm, moderately densely scabrous, truncate, obtuse, or acute, lacerate to entire;
innovation blades to 20 cm, adaxial surfaces sparsely scabrous, glabrous or hispidulous on and between the veins;
cauline blades 1.5-2.4 mm wide, folded or flat, thin, soft, abaxial surfaces smooth, veins prominent, keel and margins scabrous, adaxial surfaces moderately scabrous over the veins, sparsely scabrous between the veins, apices narrowly prow-shaped, flag leaf blades 2.9-8.6(11) cm.
mainly extravaginal.
extra-, pseudo-, and intravaginal.
(2)3.5-15 cm, ovoid to broadly pyramidal, usually open, sparse, with 10-40(60) spikelets, proximal internodes shorter than 1.5(3) cm, with (1)2-5 branches per node;
branches 1.5-6 cm, spreading soon after emergence from the sheath, thin, sinuous, and flexuous to fairly stout and straight, terete, smooth or sparsely to infrequently moderately scabrous, with (1)2-5 spikelets, the spikelets not crowded.
(4)5.5-10.5(12.5) cm, erect, ovoid to broadly pyramidal, open, or eventually loosely contracted, sparse, with 10-40 spikelets;
nodes with 1-2 branches;
branches 2.1-4.5(7) cm, ascending, lax, angled, angles moderately to densely scabrous, less scabrous between the angles, with 1-9 spikelets.
(3.5) 4.5-8 mm, lengths to 3.5 times widths, laterally compressed, sometimes bulbiferous;
florets (2)3-6, infrequently bulb-forming;
rachilla internodes smooth or muriculate, proximal internodes glabrous or sparsely softly puberulent to long-villous.
5.3-9 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;
florets (2)3-6(7);
rachilla internodes 1-1.3 mm, visible from the sides, usually sparsely to densely, coarsely scabrous, infrequently smooth.
lanceolate to broadly lanceolate, distinctly or weakly keeled, keels usually smooth, sometimes sparsely scabrous distally, lateral veins usually moderately pronounced;
lower glumes (3)3.5-5(6) mm, 3-veined;
upper glumes 3.5-5.5(6.5) mm, nearly equaling to slightly exceeding the lowest lemmas, or distinctly shorter;
calluses glabrous or webbed, hairs sparse and short to over 1/3 – 2/3 the lemma length;
lemmas (2.7)3-6(7) mm, lanceolate to broadly lanceolate, usually strongly purple, distinctly keeled, keels, marginal veins, and lateral veins long-villous, hairs on the lateral veins sometimes shorter, lateral veins prominent, intercostal regions short-villous to softly puberulent at least near the base, glabrous elsewhere, smooth to weakly muriculate and/or usually sparsely scabrous, infrequently moderately scabrous, margins broadly hyaline, glabrous, apices acute;
palea keels usually short- to long-villous for most of their length, rarely nearly glabrous and scabrous, intercostal regions broad, usually at least sparsely softly puberulent, rarely glabrous, apices scabrous;
anthers 1.4-2.5 mm, sometimes aborted late in development.
distinctly keeled;
lower glumes (2)2.7-3.8 mm, 3-veined, upper glumes (2.3)2.9-3.9 mm;
calluses diffusely webbed, hairs 1/3-1/2 the lemma length;
lemmas (3.2)4.25-5 mm, lanceolate to narrowly lanceolate, distinctly keeled, glabrous, moderately to densely, infrequently sparsely, scabrous, lateral veins prominent, margins narrowly scarious, glabrous, apices acute to narrowly acute;
paleas 3/4 as long as to subequal to the lemmas, keels scabrous, intercostal regions scabrous;
anthers (1.4)1.75-2.6 mm, or vestigial (0.1-0.2 mm).
= 36, 42, 56, 60, 62-68, 70, ca. 72, 74-76, 78-80, 82-84, 86, 88, 99, 106.
= unknown.
Poa arctica
Poa diaboli
Poa arctica is a common circumboreal species of arctic and alpine regions, growing mainly in mesic to subhydric, acidic tundra and alpine meadows, and on rocky slopes. It extends south in the Rocky Mountains to New Mexico. The frequency of sterile anthers in plants of the high arctic suggests that P. arctica is sometimes apomictic in that region. Over most of the rest of its range, P. arctica usually develops normal anthers. This and isozyme data for populations from alpine and low arctic regions suggest sexual reproduction is common in these habitats.
The most reliable way to distinguish Poa arctica from P. pratensis (p. 522), particularly subsp. alpigena, is by the wider paleas and the presence of hairs between the palea keels. Bulbiferous forms of P. arctica differ from P. stenantha var. vivipara (p. 594) in not being glaucous, and in having rhizomes and terete, smooth panicle branches. Poa xgaspensis (p. 601) also resembles P. arctica, but it has sharply keeled, more scabrous glumes and a spikelet shape that is intermediate between P. pratensis and P. alpina (p. 518). Poa arctica forms natural hybrids with both P. pratensis and P. secunda (p. 586).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Poa diaboli, which is sequentially gynomonoecious, is endemic to upper shaly slopes, in soft coastal scrub and openings in Bishop Pine stands, in the coastal mountains of San Luis Obispo County, California. It is closely related to P. confinis (see previous), from which it differs by a suite of characters. The two species are also ecologically and geographically distinct.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Plants lacking well-developed rhizomes; anthers aborted late in development; plants of the high arctic | subsp. caespitans |
1. Plants usually with well-developed rhizomes; anthers normal or plants not of the high arctic. | → 2 |
2. Panicles erect, the branches relatively stout, fairly straight; longest branches of the lowest panicle nodes 1/4-1/2 the length of the panicles; culms wiry, usually several together; calluses glabrous or shortly webbed; paleas sometimes glabrous; plants glaucous, growing in the southern Rocky-Mountains and adjacent portions of the Intermountain region | subsp. aperta |
2. Panicles lax to erect, the branches slender, flexuous to fairly stout and straight; longest branches of the lowest panicle nodes 2/5 – 3/5 the length of the panicles; culms slender to stout, varying from solitary to several together; calluses glabrous or webbed, the hairs usually more than 1/2 as long as the lemmas; paleas pubescent; plants sometimes glaucous, widespread in distribution. | → 3 |
3. Calluses glabrous; spikelets not bulbiferous | subsp. grayana |
3. Calluses webbed, often copiously so, sometimes glabrous in bulbiferous spikelets; spikelets sometimes bulbiferous. | → 4 |
4. Spikelets (5)6-8 mm long; lemmas 4-6 mm long; blades 2-6 mm wide; rachillas usually hairy; plants primarily of the western arctic, extending to northwestern British Columbia | subsp. lanata |
4. Spikelets (3.5)4-7 mm long; lemmas (2.7)3-4.5 mm long; blades 1.5-3 mm wide; rachillas commonly glabrous; plants widespread | subsp. arctica |
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