Poa annua |
Poaceae subfam. pooideae |
|||||||||||||||||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
annual blue grass |
|
|||||||||||||||||||||||||||||||||||||
Habit | Plants usually annual, rarely surviving for a second season; not rhizomatous, sometimes stoloniferous, densely tufted. | Plants annual or perennial; sometimes matlike, sometimes cespitose, sometimes stoloniferous, sometimes rhizomatous. | ||||||||||||||||||||||||||||||||||||
Culms | 2-20(45) cm, prostrate to erect, slender; nodes terete, usually 1 exserted. |
usually hollow, sometimes solid. |
||||||||||||||||||||||||||||||||||||
Sheaths | closed for about 1/3 their length, terete or weakly compressed, smooth; ligules 0.5-3(5) mm, smooth, glabrous, decurrent, obtuse to truncate; blades 1-10 cm long, 1-3(6) mm wide, flat or weakly folded, thin, soft, smooth, margins usually slightly scabrous, apices broadly prow-shaped. |
|||||||||||||||||||||||||||||||||||||
Basal branching | intravaginal, innovations common, similar to the culms. |
|||||||||||||||||||||||||||||||||||||
Leaves | distichous; sheaths usually open to the base, varying to closed for nearly their full length; auricles present or absent; abaxial ligules absent; adaxial ligules scarious or membranous, sometimes puberulent or scabridulous, usually not ciliate, cilia sometimes shorter than the base; pseudopetioles rarely present; blades usually linear, sometimes broadly so, venation parallel; cross sections non-Kranz, mesophyll nonradiate, adaxial palisade layer absent, fusoid and arm cells usually absent; midribs usually simple; adaxial bulliform cells present; stomates with parallel-sided subsidiary cells; epidermes usually lacking bicellular microhairs, sometimes with unicellular microhairs, papillae usually absent, when present, rarely more than 1 per cell. |
|||||||||||||||||||||||||||||||||||||
Panicles | 1-7(10) cm, lengths 1.2-1.6 times widths, erect; nodes with 1-2(3) branches; branches ascending to spreading or reflexed, straight, terete, smooth, with crowded or loosely arranged spikelets. |
|||||||||||||||||||||||||||||||||||||
Inflorescences | usually terminal, panicles, spikes, or racemes, usually ebracteate; disarticulation usually below the florets, sometimes below the glumes, at the rachis nodes, or at the inflorescence bases. |
|||||||||||||||||||||||||||||||||||||
Spikelets | 3-5 mm, laterally compressed; florets 2-6; rachilla internodes smooth, glabrous, concealed or exposed, distal internodes less than 1/2(3/4) the length of the distal lemma. |
usually bisexual, infrequently unisexual or mixed, usually laterally compressed or not compressed, occasionally dorsally compressed, with 1-30 sexual florets, distal floret(s) often reduced, infrequently spikelets with 1-2 reduced or staminate basal florets and a single terminal sexual floret. |
||||||||||||||||||||||||||||||||||||
Glumes | smooth, distinctly keeled, keels smooth; lower glumes 1-veined; upper glumes shorter than or subequal to the lowest lemma; calluses glabrous; lemmas 2.5-4 mm, lanceolate, distinctly keeled, smooth throughout, the keels, marginal veins, and, usually, lateral veins crisply puberulent to long-villous, rarely glabrous throughout, lateral veins prominent, intercostal regions glabrous, margins smooth, glabrous, apices obtuse to acute; palea keels smooth, usually short- to long-villous, rarely glabrous; anthers 0.6-1.1 mm, oblong prior to dehiscence, those of the upper 1-2 florets usually vestigial. |
usually 2, upper or lower glumes sometimes absent, rarely both glumes absent; lemmas without uncinate hairs, awned or not, awns single, basal to apical; paleas usually well-developed, sometimes reduced or absent; lodicules 2(3), usually lanceolate and broadly membranous distally, rarely truncate and fleshy, usually not veined or obscurely veined, sometimes distinctly veined, sometimes ciliate; anthers (1, 2)3; ovaries glabrous or sometimes hairy distally, sometimes with an apical appendage; haustorial synergids absent; styles (1)2 (-4), bases close together, sometimes fused. |
||||||||||||||||||||||||||||||||||||
Caryopses | hila linear, elliptic, ovate, or punctate; endosperm usually hard, sometimes soft or liquid, with or without lipids, starch grains compound or simple; embryos less than 1/2 the length of the caryopses; epiblasts usually present; scutellar cleft usually absent; mesocotyl internode usually absent; embryonic leaf margins overlapping, x = 7, 10. |
|||||||||||||||||||||||||||||||||||||
2n | = 28. |
|||||||||||||||||||||||||||||||||||||
Poa annua |
Poaceae subfam. pooideae |
|||||||||||||||||||||||||||||||||||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; PR; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Greenland
|
|||||||||||||||||||||||||||||||||||||
Discussion | Poa annua is one of the world's most widespread weeds. It thrives in anthropomorphic habitats outside of the arctic. A native of Eurasia, it is now well established throughout most of the Flora region. Poa annua is a gynomonoecious tetraploid (possibly rarely polyhaploid), and is thought to have arisen from hybridization between P. infirma (see next) and P. supina (p. 521) (Tutin 1952). It is similar to P. infirma, differing in having larger anthers. It differs from P. chapmaniana (p. 534) in having glabrous calluses and three larger anthers, rather than one. Forms with glabrous lemmas occur sporadically within populations. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The subfamily Pooideae includes approximately 3300 species, making it the largest subfamily in the Poaceae. It reaches its greatest diversity in cool temperate and boreal regions, extending across the tropics only in high mountains. The circumscription and relationships of tribes within the Pooideae are unsettled (see, for example, Catalan et al. 1997, 2004; Soreng and Davis 1998). In this flora, some previously recognized tribes have been combined with the Poeae. Recognition of some of these as subtribes is well supported; among these is the Hainardieae Greuter (which, at the subtribal level, is called the Parapholiinae Caro). Members of other traditional tribal groupings, such as the Aveneae Dumort., appear to be widely dispersed within the Poeae sensu lato. Further work will probably support the division of the expanded Poeae into additional tribes; there is as yet no clear indication as to what the boundaries of such tribes should be. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||||||||||||||||||||||||||
Key |
|
|||||||||||||||||||||||||||||||||||||
Source | FNA vol. 24, p. 519. | FNA vol. 24, p. 57. | ||||||||||||||||||||||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Micrantherae | Poaceae | ||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||||||||||
Synonyms | P. annua var. reptans, P. annua var. aquatica | |||||||||||||||||||||||||||||||||||||
Name authority | L. | Benth. | ||||||||||||||||||||||||||||||||||||
Web links |
|