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annual blue grass

Howell's blue grass

Habit Plants usually annual, rarely surviving for a second season; not rhizomatous, sometimes stoloniferous, densely tufted. Plants usually annual, rarely longer-lived; densely tufted, tuft bases narrow; not stoloniferous, not rhizomatous.
Culms

2-20(45) cm, prostrate to erect, slender;

nodes terete, usually 1 exserted.

(10) 25-80(120) cm tall, 0.4-1.75 mm thick, usually erect;

nodes terete, usually 1-2 exserted.

Sheaths

closed for about 1/3 their length, terete or weakly compressed, smooth;

ligules 0.5-3(5) mm, smooth, glabrous, decurrent, obtuse to truncate;

blades 1-10 cm long, 1-3(6) mm wide, flat or weakly folded, thin, soft, smooth, margins usually slightly scabrous, apices broadly prow-shaped.

closed for 1/2 - 7/8 their length, usually weakly compressed and keeled, usually scabrous, rarely smooth;

ligules 1.5-5(10) mm, smooth or scabrous, acute;

blades 1-7(10) mm wide, flat, lax, soft, finely scabrous, apices narrowly prow-shaped, cauline blades 2-10 cm.

Basal branching

intravaginal, innovations common, similar to the culms.

intravaginal.

Panicles

1-7(10) cm, lengths 1.2-1.6 times widths, erect;

nodes with 1-2(3) branches;

branches ascending to spreading or reflexed, straight, terete, smooth, with crowded or loosely arranged spikelets.

10-25(30) cm, erect, eventually open, with (1)3-5(7) branches per node;

branches eventually spreading or reflexed, fairly straight, angled, angles usually moderately to densely scabrous, rarely sparsely scabrous.

Spikelets

3-5 mm, laterally compressed;

florets 2-6;

rachilla internodes smooth, glabrous, concealed or exposed, distal internodes less than 1/2(3/4) the length of the distal lemma.

(2)4-6 mm, laterally compressed, with 2-5 florets;

rachilla internodes about 1 mm, smooth, usually softly puberulent, infrequently glabrous.

Glumes

smooth, distinctly keeled, keels smooth;

lower glumes 1-veined;

upper glumes shorter than or subequal to the lowest lemma;

calluses glabrous;

lemmas 2.5-4 mm, lanceolate, distinctly keeled, smooth throughout, the keels, marginal veins, and, usually, lateral veins crisply puberulent to long-villous, rarely glabrous throughout, lateral veins prominent, intercostal regions glabrous, margins smooth, glabrous, apices obtuse to acute;

palea keels smooth, usually short- to long-villous, rarely glabrous;

anthers 0.6-1.1 mm, oblong prior to dehiscence, those of the upper 1-2 florets usually vestigial.

slightly unequal, lanceolate, distinctly keeled, keels and sometimes the lateral veins sparsely to moderately scabrous;

lower glumes 1-3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses of some or all florets sparsely webbed;

lemmas 2.5-3.5 mm, lanceolate to narrowly lanceolate, distinctly keeled, crisply puberulent proximally, hairs evenly distributed, finely scabrous distally, lateral veins obscure to prominent, margins narrowly hyaline, glabrous, apices narrowly acute, infrequently anthocyanic;

palea keels sparsely scabrous, glabrous or softly puberulent at midlength, intercostal regions usually softly puberulent;

anthers 3, 0.2-1 mm.

2n

= 28.

= unknown.

Poa annua

Poa howellii

Distribution
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; PR; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa annua is one of the world's most widespread weeds. It thrives in anthropomorphic habitats outside of the arctic. A native of Eurasia, it is now well established throughout most of the Flora region.

Poa annua is a gynomonoecious tetraploid (possibly rarely polyhaploid), and is thought to have arisen from hybridization between P. infirma (see next) and P. supina (p. 521) (Tutin 1952). It is similar to P. infirma, differing in having larger anthers. It differs from P. chapmaniana (p. 534) in having glabrous calluses and three larger anthers, rather than one. Forms with glabrous lemmas occur sporadically within populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa howellii grows primarily on rocky banks and wooded slopes, from the coastal ranges of southern British Columbia to southern California. It differs from P. bolanderi (see above) in having puberulent, rather than smooth or scabrous, lemmas, and in growing at lower elevations, mostly from near sea level to 1000 m.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 519. FNA vol. 24, p. 534.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Micrantherae Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. annua var. reptans, P. annua var. aquatica P. bolanderi var. howellii
Name authority L. Vasey & Scribn.
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