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annual semaphoregrass, California semaphoregrass, semaphore grass

nodding false semaphore grass, nodding semaphore grass

Habit Plants annual or perennial; cespitose, with many innovations, not rhizomatous, or with poorly developed rhizomes. Plants perennial; not cespitose, rhizomatous.
Culms

15-95 cm, erect or geniculate at the base, sometimes rooting at the lower nodes;

nodes 3-6.

(85)100-150 cm, erect.

Sheaths

glabrous;

ligules 2-6 mm;

blades 3-29 cm long, 3-8 mm wide, adaxial surfaces slightly scabridulous.

glabrous or pubescent, sometimes scabridulous;

ligules 2-7 mm;

blades 10-40 cm long, 5-14 mm wide, apices acute or acuminate and sharply mucronate, flag leaves often reduced to spinose tips.

Racemes

8-35 cm, with 6-13 spikelets;

internodes 1-6 cm;

pedicels 1-5.5(9) mm, ascending to spreading.

(10)20-35 cm, with 6-14 spikelets;

internodes 2-5.6(8) cm;

pedicels 2-3 mm long, reflexed at maturity.

Spikelets

15-60 mm, ascending to spreading, with 7-20(30) florets, lower florets bisexual, upper florets pistillate, terminal florets sterile.

(20)25-28 mm, pendent, with 7-14 florets, usually all but the terminal floret bisexual.

Glumes

lanceolate to broadly ovate;

lower glumes 1-4.5 mm;

upper glumes 2-6.5 mm, 1-3-veined;

rachilla internodes 1-2.8 mm long, 0.2-0.5 mm thick, with a glandular swelling at the base;

lemmas 4.5-7.5 mm, 7(9)-veined, veins distinct, often prominent, apices truncate or emarginate, unawned or awned, awns to 11 mm;

paleal keels scabridulous, each with a 0.5-2.5 mm triangular, winglike appendage, often denticulate beyond the appendage;

anthers 2.5-4 mm.

frequently scabrous, apices obtuse, sometimes erosely notched;

lower glumes 3-6 mm;

upper glumes 4-7(8.3) mm, 3-veined;

rachilla internodes 2-3 mm long, about 0.1 mm thick, bases sometimes weakly differentiated into a glandular swelling;

lemmas 8-10 mm, faintly 7(9)-veined, lateral veins occasionally prominent, apices truncate, sometimes toothed, awned, awns (5)9-20 mm;

paleal keels each with a 0.2-0.6(1) mm triangular appendage;

anthers 3.5-4 mm.

Caryopses

2.5-3.1 mm.

4.5-6 mm.

2n

=32, 36.

Pleuropogon californicus

Pleuropogon refractus

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Pleuropogon refractus grows in wet meadows, riverbanks, and shady places, from sea level to about 1000 m. Its range extends from British Columbia south to California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants annual or facultative perennials; lemmas usually with awns 5-11 mm long, rarely unawned; paleal appendages 0.5-2.5 mm long; spikelets 15-30 mm long
var. californicus
1. Plants perennial; lemmas unawned, sometimes mucronate, mucros to 1.5 mm long; paleal appendages 0.5-1 mm long; spikelets 25-60 mm long
var. davyi
Source FNA vol. 24. FNA vol. 24.
Parent taxa Poaceae > subfam. Pooideae > tribe Meliceae > Pleuropogon Poaceae > subfam. Pooideae > tribe Meliceae > Pleuropogon
Sibling taxa
P. hooverianus, P. oregonus, P. refractus, P. sabinei
P. californicus, P. hooverianus, P. oregonus, P. sabinei
Subordinate taxa
P. californicus var. californicus, P. californicus var. davyi
Name authority (Nees) Benth. ex Vasey (A. Gray) Benth. ex Vasey
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