Platanthera huronensis
Platanthera orbiculata
great lakes rein orchid, green bog orchid, Huron green orchid, Lake Huron bog green orchid, northern green bog-orchid
large round-leaf orchid, large round-leaf rein orchid, lesser roundleaf orchid, round-leaf bog-orchid, round-leaf orchid
few–several, ascending, scattered along stem, gradually reduced to bracts distally;
blade oblong to linear-lanceolate, 5–30 × 0.6–7 cm.
2, in subequal basal pair lying on ground;
bracts (very rarely, 0–)1–6, scattered along stem;
blade broadly elliptic to orbiculate or oblate, 5–21 × 3–22 cm.
very lax to very dense.
lax to dense.
resupinate, not showy but sometimes conspicuous, whitish green;
corolla often whiter than calyx;
lateral sepals spreading to slightly reflexed;
petals ovate- to lance-falcate, margins entire;
lip descending or apex adhering to dorsal sepal and petal apices, lanceolate to nearly linear, without basal thickening, 5–12 × 2–4 mm, base slightly to rather markedly rounded-dilated, margins entire;
spur slenderly cylindric to clavate, 4–12 mm, apex usually slenderly tapered;
rostellum lobes divergent, directed downward, very small, obscure, rounded;
pollinaria straight;
pollinia remaining enclosed in anther sacs;
viscidia oblong;
ovary rather slender to stout, mostly 6–15 mm.
resupinate, rather showy;
calyx mostly greenish white;
corolla mostly white;
lateral sepals reflexed to somewhat spreading;
petals lanceolate-falcate, margins entire;
lip descending to somewhat reflexed, linear-oblong to linear-lanceolate, without basal thickening, 7–17 × 1–2.5 mm, margins entire;
spur slenderly clavate, 14–27 mm;
rostellum lobes directed strongly forward, wide-spreading, angular;
pollinaria nearly straight, 3–4.5(–4.8, very rarely) mm;
pollinia remaining enclosed in anther sacs;
viscidia orbiculate;
ovary slender to rather stout, mostly 10–26 mm.
= 84.
= 42.
Platanthera huronensis
Platanthera orbiculata
Northwestern plants commonly treated as Platanthera hyperborea var. viridiflora (Chamisso) Kitamura (note Kitamura’s priority over Luer) are P. huronensis; Chamisso’s name furthermore is synonymous with P. stricta. Aleutian and coastal Alaskan plants are often short, stout, and broad-leaved, and they have incorrectly been referred to 9. P. convallariifolia.
Platanthera huronensis as here delimited does not auto-pollinate in the manner of P. aquilonis. Occasional plants and populations that may be referable to P. huronensis, however, exhibit the movement of pollinia typical of P. aquilonis. These plants might reflect infraspecific variation within an allotetraploid species, result from hybridization, or constitute a distinct taxon. The relationship of some of these plants to P. hyperborea needs study.
Platanthera huronensis is typically intensely fragrant with the sweet, pungent scent of some related species.
Platanthera huronensis is known to hybridize with P. dilatata; it may hybridize with other species as well. Although hybrids of P. dilatata and P. aquilonis may occur, the name traditionally used for them, P. ×media (Rydberg) Luer is a synonym of P. huronensis. See notes under 10. P. aquilonis and 8. P. hyperborea.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
The key will permit determination of most specimens of Platanthera orbiculata and P. macrophylla. More precise discrimination of ambiguous specimens is possible by use of the formula from A. H. Reddoch and J. M. Reddoch (1993): spur length + (2 × pollinarium length), which for P. orbiculata is less than 38 and for P. macrophylla is equal to or greater than 38.
Considerable variation in size and shape of leaves occurs, and although to some extent regional in nature, intergradation is complete; recognition of infraspecific taxa is unwarranted. A few collections from isolated areas on the Pacific Coast of Canada are noteworthy, however. Those are small, few-flowered plants with rather narrow leaves borne alternately or suboppositely toward the base of the stem, as in some Asiatic species. They are in some respects very similar to Platanthera freynii Kränzlin, an Asiatic species distinguished primarily by its abruptly narrowed petals, in contrast to the generally broader, but variable, petals in North American plants. These western plants warrant further study to establish their identity and to elucidate relationships between North American and Asiatic species.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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