Platanthera clavellata
Platanthera convallariifolia
little club-spur bog-orchid, small green wood orchid
1(–2), on basal half of stem, ascending to spreading, usually abruptly reduced to bracts distally;
blade oblanceolate to oblong, linear-oblong, or elliptic, 3–19 × 0.8–3.5 cm.
few–several, ascending to rather wide spreading, scattered along stem or commonly loosely clustered on distal portion, gradually reduced to bracts distally;
blade elliptic-lanceolate to elliptic-oblong, 4.5–12 × 0.7–2.5 cm.
moderately dense.
moderately dense.
resupinate, often incompletely so and held at angle, not showy, rather inconspicuous pale green to dull yellowish green;
lateral sepals porrect;
petals ovate to obovate;
lip oblong, obscurely 3-lobed, without basal thickening, 3–7 × 3–4 mm, margins sometimes dentate-lacerate, apex truncate;
spur clavate, 7–13 mm;
rostellum lobes directed downward, very short, truncate;
pollinaria straight or slightly curved laterally;
pollinia fragmenting, pollen masses trailing down onto stigma;
viscidia linear to linear lanceolate;
ovary rather stout, 6–11 mm.
resupinate, not showy, sometimes conspicuous, whitish green, perhaps more yellowish on lip;
lateral sepals spreading;
petals ovate-falcate, margins entire;
lip descending, linear to lance-linear, without basal thickening, 4–9 × 1–2 mm, margins entire;
spur slenderly to markedly clavate, rarely more slender, 4–8 mm, apex obtuse;
rostellum lobes widely divergent, directed downward, slightly angular, very small, obscure;
pollinaria straight;
pollinia remaining enclosed in anther sacs but apparently often fragmenting;
viscidia oblong to oblong-spatulate or elliptic-suborbiculate;
ovary slender to stout, mostly 6–10 mm.
= 42.
Platanthera clavellata
Platanthera convallariifolia
In common with Platanthera nivea and P. integra, the column of P. clavellata bears a series of lateral projections that are directed forward; the upper pair is elaborately adorned and may be glandular. It appears that this species is inappropriately placed in Platanthera. See note under 30. P. nivea.
The broader leaves cited as distinguishing Platanthera clavellata var. ophioglossoides are more prevalent in the North, but occur throughout the range of the species. In some areas populations commonly display a complete range of leaf shape; this feature alone is of no taxonomic significance.
The unusual and infrequent hybrid Platanthera blephariglottis × P. clavellata is P. ×vossii Case.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Platanthera convallariifolia is commonly reported from the Aleutians and adjacent coastal Alaska based on short, very broad-leaved plants with rather large flowers in dense, thick inflorescences. Those plants, however, apparently represent robust, maritime phenotypes of P. huronensis or in some cases P. stricta or P. dilatata comparable to those seen in Piperia elegans and Spiranthes romanzoffiana. In fact, typical Asiatic Platanthera convallariifolia is a slender-leaved plant entirely comparable in habit to P. huronensis. The two species are very similar, and they share a tetraploid chromosome number. Typical Asiatic P. convallariifolia is characterized by a linear lip and clavate spur about 2/3 to nearly the length of the lip. The lip occasionally varies to slightly lanceolate and the spur to somewhat saccate. The column appears somewhat broader than in P. huronensis, and viscidia also are apparently rather broad. A few specimens from the Aleutians and Alaskan peninsula exhibit these floral characteristics. Vegetatively, however, some Alaskan material is distinctive in the position and orientation of the leaves. In floral and vegetative characters, Alaskan plants furthermore intergrade with P. huronensis. Whether Alaskan plants are truly referable to P. convallariifolia is uncertain, and the status of the Asiatic species also needs study. Both P. convallariifolia and the Alaskan plants here referred to it appear to combine features of P. huronensis and P. stricta. Platanthera convallariifolia may have arisen through hybridization of these two species, or of P. stricta and P. dilatata, thus accounting for the difficulty in identification. The description presented here is based on few Alaskan plants and is apt to be too restrictive; therefore the poor understanding of the plant makes the present delimitation and distinctions from P. huronensis rather arbitrary.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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