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Tweedy's plantain

blond plantain, desert Indian-wheat, desert plantain

Habit Perennials; caudex well developed, conspicuous, glabrous; roots taproots, thick. Annuals; roots taproots, slender.
Stems

0–20 mm.

0–30 mm, often branched.

Leaves

40–200 × 10–30 mm;

blade lanceolate-spatulate to narrowly ovate, margins entire, sometimes toothed, veins conspicuous, surfaces usually glabrous.

10–230 × 0.5–12 mm;

blade linear or narrowly elliptic, margins toothed, veins conspicuous or not, surfaces villous or lanate to sericeous.

Scapes

25–200 mm, slightly surpassing leaves, glabrous.

10–400 mm, hairy, hairs woolly, long.

Spikes

brownish or greenish, 45–250 mm, densely flowered, rachis not clearly visible between flowers;

bracts broadly ovate, 2 mm, length 0.8–1 times sepals.

grayish or brownish, 20–400 mm, densely flowered, flowers in spirals;

bracts ovate or elliptic, 1.7–4 mm, length 0.8–1.2 times sepals, apex not reached by green nerve.

Flowers

sepals 2–2.5 mm;

corolla radially symmetric, lobes spreading, 1 mm, base obtuse;

stamens 4.

sepals 1.9–3.5 mm;

corolla radially symmetric, lobes reflexed, 1.3–2.8 mm, base cuneate;

stamens 4.

Seeds

3 or 4, 2–2.3 mm.

2, 2–2.6 mm.

2n

= 24.

= 8.

Plantago tweedyi

Plantago ovata

Phenology Flowering summer. Flowering spring.
Habitat Grasslands, sagebrush steppes, montane and subalpine meadows. Sandy deserts and steppes.
Elevation 1600–4000 m. (5200–13100 ft.) 0–1800 m. (0–5900 ft.)
Distribution
from FNA
AZ; CO; ID; MT; NM; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; TX; UT; Mexico (Baja California, Sonora); Eurasia; Africa
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[BONAP county map]
Discussion

From molecular evidence, S. C. Meyers and A. Liston (2008) suggested that Plantago ovata was introduced to North America during the Pleistocene. They recognized four varieties; North American specimens can be treated as two varieties based on bract and corolla color: the inland var. fastigiata (E. Morris) S. C. Meyers & Liston (midribs of mature flower bracts green, corolla lobes without reddish brown midribs) and the coastal var. insularis (Eastwood) S. C. Meyers & Liston (midribs of mature flower bracts brown, corolla lobe midribs prominent, reddish brown). Unfortunately, these features are not easily seen on many herbarium specimens, and these taxa are not recognized here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 293. FNA vol. 17, p. 290.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. insularis, P. insularis var. fastigiata
Name authority A. Gray: in A. Gray et al., Syn. Fl. N. Amer. ed. 2, 2(1): 390. (1886) Forsskål: Fl. Aegypt.-Arab., 31. (1775)
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