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pineland plantain

blackseed plantain, plantain de Rugel, Rugel's plantain

Habit Perennials; caudex well developed, conspicuous, glabrous; roots taproots, thick. Perennials, sometimes annuals; caudex absent; roots fibrous, thick.
Stems

0–10 mm.

0–20 mm.

Leaves

50–300 × 10–30 mm;

blade lanceolate, margins entire, sometimes toothed, veins conspicuous, laterals branching from base, surfaces sparsely hairy.

20–150 × 10–120 mm;

petiole to 200 mm;

blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute.

Scapes

75–150 mm, sparsely hairy.

50–250 mm, glabrous or hirsute.

Spikes

brownish or greenish, 100–450 mm, loosely flowered, rachis visible between flowers;

bracts ovate, 1 mm, length 0.5–0.6 times sepals.

brownish or greenish, 50–300 mm, densely or loosely flowered;

bracts narrowly lanceolate, 2 mm, length 1–1.2 times sepals.

Flowers

sepals 2 mm;

corolla radially symmetric, lobes spreading, 1 mm, base obtuse;

stamens 4.

sepals 1.5–2 mm;

corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse;

stamens 4.

Fruits

4–6(–8) mm, dehiscing proximal to middle.

Seeds

2, 2 mm.

4 or 5(–8), 1.5–2 mm.

2n

= 24.

= 24.

Plantago sparsiflora

Plantago rugelii

Phenology Flowering summer. Flowering summer.
Habitat Moist, sandy soils, open, undisturbed pine woods. Open woods, meadows, pastures, waste places.
Elevation 0–200 m. (0–700 ft.) 0–2000 m. (0–6600 ft.)
Distribution
from FNA
FL; GA; NC; SC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Populations of Plantago sparsiflora occur mainly along the Atlantic coast from Columbia County, North Carolina, south to Volusia County, Florida.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants of Plantago major and P. rugelii are morphologically indistinguishable when young; they may be differentiated with certainty only at fruiting stage. DNA sequence data (A. Shipunov et al. 2014) confirm that the two species are distinct: P. rugelii is different from P. major by 11 substitutions in the ITS2 sequence.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 292. FNA vol. 17, p. 292.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Name authority Michaux: Fl. Bor.-Amer. 1: 94. (1803) Decaisne: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(1): 700. (1852)
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