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evergreen plantain

blond plantain, desert Indian-wheat, desert plantain

Habit Perennials, sometimes woody; roots taproots, slender. Annuals; roots taproots, slender.
Stems

100–400 mm, freely branched.

0–30 mm, often branched.

Leaves

cauline, opposite, 30–60 × 0.75–1 mm;

blade linear to linear-lanceolate, margins entire, veins inconspicuous, surfaces hairy.

10–230 × 0.5–12 mm;

blade linear or narrowly elliptic, margins toothed, veins conspicuous or not, surfaces villous or lanate to sericeous.

Scapes

30–80 mm, hairy.

10–400 mm, hairy, hairs woolly, long.

Spikes

greenish or brownish, 40–85 mm, densely flowered;

bracts broadly ovate, 5–6 mm, lengths equal to sepals.

grayish or brownish, 20–400 mm, densely flowered, flowers in spirals;

bracts ovate or elliptic, 1.7–4 mm, length 0.8–1.2 times sepals, apex not reached by green nerve.

Flowers

sepals 5–6 mm;

corolla radially symmetric, lobes reflexed, 3–3.5 mm, base obtuse;

stamens 4.

sepals 1.9–3.5 mm;

corolla radially symmetric, lobes reflexed, 1.3–2.8 mm, base cuneate;

stamens 4.

Seeds

1 or 2, 2 mm.

2, 2–2.6 mm.

2n

= 12.

= 8.

Plantago sempervirens

Plantago ovata

Phenology Flowering summer–fall. Flowering spring.
Habitat Disturbed habitats. Sandy deserts and steppes.
Elevation 0–200 m. (0–700 ft.) 0–1800 m. (0–5900 ft.)
Distribution
from FNA
CA; s Europe [Introduced in North America]
from FNA
AZ; CA; NV; TX; UT; Mexico (Baja California, Sonora); Eurasia; Africa
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[BONAP county map]
Discussion

Plantago sempervirens is known from San Diego County; one specimen (JEPS) was collected in 2008 in Torrey Pines State Reserve.

Plantago cynops Linnaeus (1762, not 1753), a rejected name, has been misapplied to P. sempervirens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

From molecular evidence, S. C. Meyers and A. Liston (2008) suggested that Plantago ovata was introduced to North America during the Pleistocene. They recognized four varieties; North American specimens can be treated as two varieties based on bract and corolla color: the inland var. fastigiata (E. Morris) S. C. Meyers & Liston (midribs of mature flower bracts green, corolla lobes without reddish brown midribs) and the coastal var. insularis (Eastwood) S. C. Meyers & Liston (midribs of mature flower bracts brown, corolla lobe midribs prominent, reddish brown). Unfortunately, these features are not easily seen on many herbarium specimens, and these taxa are not recognized here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 292. FNA vol. 17, p. 290.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. insularis, P. insularis var. fastigiata
Name authority Crantz: Inst. Rei Herb. 2: 331. (1766) Forsskål: Fl. Aegypt.-Arab., 31. (1775)
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