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blackseed plantain, plantain de Rugel, Rugel's plantain

dwarf plantain, little plantain, woolly indianwheat, woolly plantain

Habit Perennials, sometimes annuals; caudex absent; roots fibrous, thick. Annuals; roots taproots, slender.
Stems

0–20 mm.

0–5 mm.

Leaves

20–150 × 10–120 mm;

petiole to 200 mm;

blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute.

20–70 × 1–2 mm;

blade linear, margins entire, sometimes toothed, veins conspicuous or not, surfaces hairy or glabrous.

Scapes

50–250 mm, glabrous or hirsute.

15–60 mm, hairy, sometimes glabrous.

Spikes

brownish or greenish, 50–300 mm, densely or loosely flowered;

bracts narrowly lanceolate, 2 mm, length 1–1.2 times sepals.

greenish or brownish, 20–100 mm, loosely or densely flowered;

bracts triangular-ovate, 1.5–2 mm, length 0.9–1.1 times sepals.

Flowers

sepals 1.5–2 mm;

corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse;

stamens 4.

sepals 1.5–2 mm;

corolla radially symmetric, lobes erect, forming a beak, 0.5 mm, base obtuse;

stamens 2.

Fruits

4–6(–8) mm, dehiscing proximal to middle.

Seeds

4 or 5(–8), 1.5–2 mm.

4, 0.8–1.3 mm.

2n

= 24.

= 12.

Plantago rugelii

Plantago pusilla

Phenology Flowering summer. Flowering spring–early summer.
Habitat Open woods, meadows, pastures, waste places. Dry to moist, sandy, alluvial soils.
Elevation 0–2000 m. (0–6600 ft.) 0–200 m. (0–700 ft.)
Distribution
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; CT; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; MO; MS; NC; NJ; NY; OK; OR; PA; RI; SC; TN; TX; VA; WA
[WildflowerSearch map]
Discussion

Plants of Plantago major and P. rugelii are morphologically indistinguishable when young; they may be differentiated with certainty only at fruiting stage. DNA sequence data (A. Shipunov et al. 2014) confirm that the two species are distinct: P. rugelii is different from P. major by 11 substitutions in the ITS2 sequence.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plantago pusilla is considered to be introduced in Oregon and Washington, and possibly in California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 292. FNA vol. 17, p. 291.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. hybrida, P. pusilla var. major
Name authority Decaisne: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(1): 700. (1852) Nuttall: Gen. N. Amer. Pl. 1: 100. (1818)
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