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blackseed plantain, plantain de Rugel, Rugel's plantain

Heller's plantain

Habit Perennials, sometimes annuals; caudex absent; roots fibrous, thick. Annuals; roots taproots, slender.
Stems

0–20 mm.

0–12 mm.

Leaves

20–150 × 10–120 mm;

petiole to 200 mm;

blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute.

50–130 × 3–7 mm;

blade linear, margins entire, veins conspicuous or not, surfaces villous, adaxial rarely glabrous.

Scapes

50–250 mm, glabrous or hirsute.

80–250 mm, hairy, hairs patent, long and antrorse, short.

Spikes

brownish or greenish, 50–300 mm, densely or loosely flowered;

bracts narrowly lanceolate, 2 mm, length 1–1.2 times sepals.

greenish, whitish, or brownish, 50–120 mm, densely flowered, flowers in whorls or pairs;

bracts triangular, 2.5–8 mm, length 0.6–2.2 times sepals.

Flowers

sepals 1.5–2 mm;

corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse;

stamens 4.

sepals 3.5–4 mm;

corolla radially symmetric, lobes reflexed, 3–3.6 mm, base slightly cordate;

stamens 4, connective elongated, apex acute.

Fruits

4–6(–8) mm, dehiscing proximal to middle.

Seeds

4 or 5(–8), 1.5–2 mm.

2, 3.2–3.8 mm.

2n

= 24.

= 20.

Plantago rugelii

Plantago helleri

Phenology Flowering summer. Flowering spring.
Habitat Open woods, meadows, pastures, waste places. Dry slopes and flats on limestone.
Elevation 0–2000 m. (0–6600 ft.) 100–1700 m. (300–5600 ft.)
Distribution
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
NM; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Plants of Plantago major and P. rugelii are morphologically indistinguishable when young; they may be differentiated with certainty only at fruiting stage. DNA sequence data (A. Shipunov et al. 2014) confirm that the two species are distinct: P. rugelii is different from P. major by 11 substitutions in the ITS2 sequence.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Populations of Plantago helleri are concentrated in central Texas, especially in the Edwards Plateau and Trans-Pecos. The species also occurs in Eddy and Lincoln counties, New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 292. FNA vol. 17, p. 287.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Name authority Decaisne: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(1): 700. (1852) Small: Bull. New York Bot. Gard. 1: 288. (1899)
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