Plantago rugelii |
Plantago helleri |
|
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blackseed plantain, plantain de Rugel, Rugel's plantain |
Heller's plantain |
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Habit | Perennials, sometimes annuals; caudex absent; roots fibrous, thick. | Annuals; roots taproots, slender. |
Stems | 0–20 mm. |
0–12 mm. |
Leaves | 20–150 × 10–120 mm; petiole to 200 mm; blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute. |
50–130 × 3–7 mm; blade linear, margins entire, veins conspicuous or not, surfaces villous, adaxial rarely glabrous. |
Scapes | 50–250 mm, glabrous or hirsute. |
80–250 mm, hairy, hairs patent, long and antrorse, short. |
Spikes | brownish or greenish, 50–300 mm, densely or loosely flowered; bracts narrowly lanceolate, 2 mm, length 1–1.2 times sepals. |
greenish, whitish, or brownish, 50–120 mm, densely flowered, flowers in whorls or pairs; bracts triangular, 2.5–8 mm, length 0.6–2.2 times sepals. |
Flowers | sepals 1.5–2 mm; corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse; stamens 4. |
sepals 3.5–4 mm; corolla radially symmetric, lobes reflexed, 3–3.6 mm, base slightly cordate; stamens 4, connective elongated, apex acute. |
Fruits | 4–6(–8) mm, dehiscing proximal to middle. |
|
Seeds | 4 or 5(–8), 1.5–2 mm. |
2, 3.2–3.8 mm. |
2n | = 24. |
= 20. |
Plantago rugelii |
Plantago helleri |
|
Phenology | Flowering summer. | Flowering spring. |
Habitat | Open woods, meadows, pastures, waste places. | Dry slopes and flats on limestone. |
Elevation | 0–2000 m. (0–6600 ft.) | 100–1700 m. (300–5600 ft.) |
Distribution |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
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NM; TX
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Discussion | Plants of Plantago major and P. rugelii are morphologically indistinguishable when young; they may be differentiated with certainty only at fruiting stage. DNA sequence data (A. Shipunov et al. 2014) confirm that the two species are distinct: P. rugelii is different from P. major by 11 substitutions in the ITS2 sequence. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Populations of Plantago helleri are concentrated in central Texas, especially in the Edwards Plateau and Trans-Pecos. The species also occurs in Eddy and Lincoln counties, New Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 292. | FNA vol. 17, p. 287. |
Parent taxa | Plantaginaceae > Plantago | Plantaginaceae > Plantago |
Sibling taxa | ||
Name authority | Decaisne: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(1): 700. (1852) | Small: Bull. New York Bot. Gard. 1: 288. (1899) |
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