Plantago rugelii |
Plantago canescens |
|
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blackseed plantain, plantain de Rugel, Rugel's plantain |
arctic plantain, gray-pubescent plantain, Siberian plantain |
|
Habit | Perennials, sometimes annuals; caudex absent; roots fibrous, thick. | Perennials; caudex usually woolly; roots taproots, thick. |
Stems | 0–20 mm. |
0–20 mm. |
Leaves | 20–150 × 10–120 mm; petiole to 200 mm; blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute. |
ascending, 180–250 × 6–20 mm; blade linear to lanceolate or oblanceolate, margins entire, rarely toothed, veins conspicuous, surfaces glabrate or hairy, hairs 1 mm. |
Scapes | 50–250 mm, glabrous or hirsute. |
50–230 mm, not groove-angled, hairy or glabrous. |
Spikes | brownish or greenish, 50–300 mm, densely or loosely flowered; bracts narrowly lanceolate, 2 mm, length 1–1.2 times sepals. |
grayish or whitish, 80–350 mm, usually densely flowered, shiny; corolla lobes of neighboring flowers often overlapping; bracts broadly ovate, 1.8–2 mm, length 0.9–1 times sepals. |
Flowers | sepals 1.5–2 mm; corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse; stamens 4. |
sepals 2 mm, adaxial 2 nearly distinct; corolla radially symmetric, lobes reflexed, 2 mm, base obtuse; stamens 4. |
Fruits | 4–6(–8) mm, dehiscing proximal to middle. |
|
Seeds | 4 or 5(–8), 1.5–2 mm. |
3–7, 1–1.8 mm. |
2n | = 24. |
= 12. |
Plantago rugelii |
Plantago canescens |
|
Phenology | Flowering summer. | Flowering summer. |
Habitat | Open woods, meadows, pastures, waste places. | Grassy, gravelly, and rocky slopes, cliffs. |
Elevation | 0–2000 m. (0–6600 ft.) | 0–2000 m. (0–6600 ft.) |
Distribution |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
|
AK; MT; AB; BC; NT; NU; YT; Asia |
Discussion | Plants of Plantago major and P. rugelii are morphologically indistinguishable when young; they may be differentiated with certainty only at fruiting stage. DNA sequence data (A. Shipunov et al. 2014) confirm that the two species are distinct: P. rugelii is different from P. major by 11 substitutions in the ITS2 sequence. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
N. N. Tzvelev (1983) recognized six subspecies (including two in North America) within Plantago canescens; North American material is not segregated as such here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 292. | FNA vol. 17, p. 285. |
Parent taxa | Plantaginaceae > Plantago | Plantaginaceae > Plantago |
Sibling taxa | ||
Synonyms | P. septata | |
Name authority | Decaisne: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(1): 700. (1852) | Adams: Nouv. Mém. Soc. Imp. Naturalistes Moscou 9: 233, plate 13, fig. 1. (1834) |
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