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dwarf plantain, little plantain, woolly indianwheat, woolly plantain

blackseed plantain, plantain de Rugel, Rugel's plantain

Habit Annuals; roots taproots, slender. Perennials, sometimes annuals; caudex absent; roots fibrous, thick.
Stems

0–5 mm.

0–20 mm.

Leaves

20–70 × 1–2 mm;

blade linear, margins entire, sometimes toothed, veins conspicuous or not, surfaces hairy or glabrous.

20–150 × 10–120 mm;

petiole to 200 mm;

blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute.

Scapes

15–60 mm, hairy, sometimes glabrous.

50–250 mm, glabrous or hirsute.

Spikes

greenish or brownish, 20–100 mm, loosely or densely flowered;

bracts triangular-ovate, 1.5–2 mm, length 0.9–1.1 times sepals.

brownish or greenish, 50–300 mm, densely or loosely flowered;

bracts narrowly lanceolate, 2 mm, length 1–1.2 times sepals.

Flowers

sepals 1.5–2 mm;

corolla radially symmetric, lobes erect, forming a beak, 0.5 mm, base obtuse;

stamens 2.

sepals 1.5–2 mm;

corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse;

stamens 4.

Fruits

4–6(–8) mm, dehiscing proximal to middle.

Seeds

4, 0.8–1.3 mm.

4 or 5(–8), 1.5–2 mm.

2n

= 12.

= 24.

Plantago pusilla

Plantago rugelii

Phenology Flowering spring–early summer. Flowering summer.
Habitat Dry to moist, sandy, alluvial soils. Open woods, meadows, pastures, waste places.
Elevation 0–200 m. (0–700 ft.) 0–2000 m. (0–6600 ft.)
Distribution
from FNA
AL; AR; CA; CT; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; MO; MS; NC; NJ; NY; OK; OR; PA; RI; SC; TN; TX; VA; WA
[WildflowerSearch map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Plantago pusilla is considered to be introduced in Oregon and Washington, and possibly in California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants of Plantago major and P. rugelii are morphologically indistinguishable when young; they may be differentiated with certainty only at fruiting stage. DNA sequence data (A. Shipunov et al. 2014) confirm that the two species are distinct: P. rugelii is different from P. major by 11 substitutions in the ITS2 sequence.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 291. FNA vol. 17, p. 292.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. hybrida, P. pusilla var. major
Name authority Nuttall: Gen. N. Amer. Pl. 1: 100. (1818) Decaisne: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(1): 700. (1852)
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