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dwarf plantain, little plantain, woolly indianwheat, woolly plantain

common plantain, great plantain, nippleseed, plantain majeur

Habit Annuals; roots taproots, slender. Perennials, sometimes annuals; caudex absent; roots fibrous, thick.
Stems

0–5 mm.

0–20 mm.

Leaves

20–70 × 1–2 mm;

blade linear, margins entire, sometimes toothed, veins conspicuous or not, surfaces hairy or glabrous.

20–150(–400) × 10–120(–170) mm;

petiole to 200 mm;

blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute.

Scapes

15–60 mm, hairy, sometimes glabrous.

50–250(–500) mm, surpassing leaves or not, glabrous or hirsute.

Spikes

greenish or brownish, 20–100 mm, loosely or densely flowered;

bracts triangular-ovate, 1.5–2 mm, length 0.9–1.1 times sepals.

brownish or greenish, (20–)50–300(–400) mm, densely flowered;

bracts lanceolate, 0.5–1 mm, length 0.3–0.7 times sepals.

Flowers

sepals 1.5–2 mm;

corolla radially symmetric, lobes erect, forming a beak, 0.5 mm, base obtuse;

stamens 2.

sepals 1.5–2 mm;

corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse;

stamens 4.

Fruits

(2–)4–5 mm, dehiscing at middle.

Seeds

4, 0.8–1.3 mm.

5–35, 0.5–1 mm.

2n

= 12.

= 12.

Plantago pusilla

Plantago major

Phenology Flowering spring–early summer. Flowering summer.
Habitat Dry to moist, sandy, alluvial soils. Roadsides, trails, stream banks, urban areas, lawns, other disturbed areas.
Elevation 0–200 m. (0–700 ft.) 0–3000 m. (0–9800 ft.)
Distribution
from FNA
AL; AR; CA; CT; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; MO; MS; NC; NJ; NY; OK; OR; PA; RI; SC; TN; TX; VA; WA
[WildflowerSearch map]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; SPM; Greenland; Eurasia [Introduced in North America; introduced also in Mexico, Central America, South America, Africa, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Plantago pusilla is considered to be introduced in Oregon and Washington, and possibly in California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies intermedia (Gilibert) Lange is often accepted by European botanists as a separate species, Plantago uliginosa F. W. Schmidt. Observations suggest that plants referable to this taxon may occur in the United States; it has been reported from the New England states (A. Haines 2011). Subspecies intermedia is distinguished by more abundant (11–35) and smaller (0.8–1 mm) seeds, ascending spikes, ovoid fruits, and elliptic or lanceolate leaf blades. Without detailed morphologic and genetic investigations of North American plants similar to the study of European plants by M. Morgan-Richards and K. Wolff (1999), it is not possible to draw any conclusions about the status and distribution of this or any other possible infraspecific taxa of P. major in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 291. FNA vol. 17, p. 289.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. hybrida, P. pusilla var. major P. major var. pachyphylla, P. major var. pilgeri, P. major var. scopulorum
Name authority Nuttall: Gen. N. Amer. Pl. 1: 100. (1818) Linnaeus: Sp. Pl. 1: 112. (1753)
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