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hoary plantain, plantain moyen

blackseed plantain, plantain de Rugel, Rugel's plantain

Habit Perennials; caudex glabrous or hairy; roots taproots, thick. Perennials, sometimes annuals; caudex absent; roots fibrous, thick.
Stems

0–20 mm.

0–20 mm.

Leaves

prostrate, sometimes ascending, 40–200 × 30–70 mm;

blade elliptic to ovate, margins entire or toothed, veins conspicuous, surfaces hairy, hairs 0.5 mm.

20–150 × 10–120 mm;

petiole to 200 mm;

blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute.

Scapes

80–300 mm, not groove-angled, hirsute.

50–250 mm, glabrous or hirsute.

Spikes

grayish or whitish, 100–400 mm, densely flowered, shiny;

corolla lobes of neighboring flowers overlapping;

bracts ovate, 1.8–2 mm, length 0.9–1 times sepals.

brownish or greenish, 50–300 mm, densely or loosely flowered;

bracts narrowly lanceolate, 2 mm, length 1–1.2 times sepals.

Flowers

sepals 2 mm, adaxial 2 nearly distinct;

corolla radially symmetric, lobes reflexed, 1.5 mm, base obtuse;

stamens 4.

sepals 1.5–2 mm;

corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse;

stamens 4.

Fruits

4–6(–8) mm, dehiscing proximal to middle.

Seeds

2–4, 2 mm.

4 or 5(–8), 1.5–2 mm.

2n

= 12, 24.

= 24.

Plantago media

Plantago rugelii

Phenology Flowering summer. Flowering summer.
Habitat Disturbed areas, neutral and basic soils. Open woods, meadows, pastures, waste places.
Elevation 0–200 m. (0–700 ft.) 0–2000 m. (0–6600 ft.)
Distribution
from FNA
CT; IL; MA; ME; MI; NH; NJ; NY; PA; RI; WI; MB; NB; ON; QC; Eurasia [Introduced in North America]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Three varieties of Plantago media have been recognized in western Europe. It is not clear how names of those varieties apply to P. media in eastern Europe and North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants of Plantago major and P. rugelii are morphologically indistinguishable when young; they may be differentiated with certainty only at fruiting stage. DNA sequence data (A. Shipunov et al. 2014) confirm that the two species are distinct: P. rugelii is different from P. major by 11 substitutions in the ITS2 sequence.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 290. FNA vol. 17, p. 292.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Name authority Linnaeus: Sp. Pl. 1: 113. (1753) Decaisne: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(1): 700. (1852)
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