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hoary plantain, plantain moyen

common plantain, great plantain, nippleseed, plantain majeur

Habit Perennials; caudex glabrous or hairy; roots taproots, thick. Perennials, sometimes annuals; caudex absent; roots fibrous, thick.
Stems

0–20 mm.

0–20 mm.

Leaves

prostrate, sometimes ascending, 40–200 × 30–70 mm;

blade elliptic to ovate, margins entire or toothed, veins conspicuous, surfaces hairy, hairs 0.5 mm.

20–150(–400) × 10–120(–170) mm;

petiole to 200 mm;

blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute.

Scapes

80–300 mm, not groove-angled, hirsute.

50–250(–500) mm, surpassing leaves or not, glabrous or hirsute.

Spikes

grayish or whitish, 100–400 mm, densely flowered, shiny;

corolla lobes of neighboring flowers overlapping;

bracts ovate, 1.8–2 mm, length 0.9–1 times sepals.

brownish or greenish, (20–)50–300(–400) mm, densely flowered;

bracts lanceolate, 0.5–1 mm, length 0.3–0.7 times sepals.

Flowers

sepals 2 mm, adaxial 2 nearly distinct;

corolla radially symmetric, lobes reflexed, 1.5 mm, base obtuse;

stamens 4.

sepals 1.5–2 mm;

corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse;

stamens 4.

Fruits

(2–)4–5 mm, dehiscing at middle.

Seeds

2–4, 2 mm.

5–35, 0.5–1 mm.

2n

= 12, 24.

= 12.

Plantago media

Plantago major

Phenology Flowering summer. Flowering summer.
Habitat Disturbed areas, neutral and basic soils. Roadsides, trails, stream banks, urban areas, lawns, other disturbed areas.
Elevation 0–200 m. (0–700 ft.) 0–3000 m. (0–9800 ft.)
Distribution
from FNA
CT; IL; MA; ME; MI; NH; NJ; NY; PA; RI; WI; MB; NB; ON; QC; Eurasia [Introduced in North America]
[BONAP county map]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; SPM; Greenland; Eurasia [Introduced in North America; introduced also in Mexico, Central America, South America, Africa, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Three varieties of Plantago media have been recognized in western Europe. It is not clear how names of those varieties apply to P. media in eastern Europe and North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies intermedia (Gilibert) Lange is often accepted by European botanists as a separate species, Plantago uliginosa F. W. Schmidt. Observations suggest that plants referable to this taxon may occur in the United States; it has been reported from the New England states (A. Haines 2011). Subspecies intermedia is distinguished by more abundant (11–35) and smaller (0.8–1 mm) seeds, ascending spikes, ovoid fruits, and elliptic or lanceolate leaf blades. Without detailed morphologic and genetic investigations of North American plants similar to the study of European plants by M. Morgan-Richards and K. Wolff (1999), it is not possible to draw any conclusions about the status and distribution of this or any other possible infraspecific taxa of P. major in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 290. FNA vol. 17, p. 289.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. major var. pachyphylla, P. major var. pilgeri, P. major var. scopulorum
Name authority Linnaeus: Sp. Pl. 1: 113. (1753) Linnaeus: Sp. Pl. 1: 112. (1753)
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