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common plantain, great plantain, nippleseed, plantain majeur

coastal plantain, Mexican plantain, tall coastal plantain

Habit Perennials, sometimes annuals; caudex absent; roots fibrous, thick. Perennials; caudex well developed, conspicuous, glabrous or hairy; roots taproots, fragile.
Stems

0–20 mm.

0–10 mm.

Leaves

20–150(–400) × 10–120(–170) mm;

petiole to 200 mm;

blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute.

60–360 × 15–65 mm;

blade elliptic to narrowly elliptic, margins toothed, veins conspicuous, surfaces pilose, rarely glabrate, adaxial surface hairs not floccose, less than 2 mm long, more than 0.03 mm wide.

Scapes

50–250(–500) mm, surpassing leaves or not, glabrous or hirsute.

55–360 mm, hairy, hairs antrorse, long.

Spikes

brownish or greenish, (20–)50–300(–400) mm, densely flowered;

bracts lanceolate, 0.5–1 mm, length 0.3–0.7 times sepals.

greenish or brownish, 110–720 mm, densely or loosely flowered;

bracts ovate, rarely triangular, 2.5–4 mm, length 0.8–1.3 times sepals.

Flowers

sepals 1.5–2 mm;

corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse;

stamens 4.

sepals 2.6–3.1 mm;

corolla radially symmetric, lobes erect, forming a beak, 2.4–2.7 mm, base obtuse;

stamens 4.

Fruits

(2–)4–5 mm, dehiscing at middle.

Seeds

5–35, 0.5–1 mm.

3, 1.8–2.5 mm.

2n

= 12.

= 48.

Plantago major

Plantago subnuda

Phenology Flowering summer. Flowering late spring–fall.
Habitat Roadsides, trails, stream banks, urban areas, lawns, other disturbed areas. Moist ground.
Elevation 0–3000 m. [0–9800 ft.] 0–300 m. [0–1000 ft.]
Distribution
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; SPM; Greenland; Eurasia [Introduced in North America; introduced also in Mexico, Central America, South America, Africa, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies intermedia (Gilibert) Lange is often accepted by European botanists as a separate species, Plantago uliginosa F. W. Schmidt. Observations suggest that plants referable to this taxon may occur in the United States; it has been reported from the New England states (A. Haines 2011). Subspecies intermedia is distinguished by more abundant (11–35) and smaller (0.8–1 mm) seeds, ascending spikes, ovoid fruits, and elliptic or lanceolate leaf blades. Without detailed morphologic and genetic investigations of North American plants similar to the study of European plants by M. Morgan-Richards and K. Wolff (1999), it is not possible to draw any conclusions about the status and distribution of this or any other possible infraspecific taxa of P. major in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plantago subnuda occurs primarily in counties along the Pacific coast from southwestern Washington to southern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 289. Treatment author: Alexey Shipunov. FNA vol. 17, p. 293. Treatment author: Alexey Shipunov.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. major var. pachyphylla, P. major var. pilgeri, P. major var. scopulorum
Name authority Linnaeus: Sp. Pl. 1: 112. (1753) Pilger: Notizbl. Königl. Bot. Gart. Berlin 5: 260. (1912)
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