Plantago major |
Plantago cordata |
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common plantain, great plantain, nippleseed, plantain majeur |
heart-leaf plantain |
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Habit | Perennials, sometimes annuals; caudex absent; roots fibrous, thick. | Perennials; caudex well developed, conspicuous, glabrous; roots fibrous, thick. |
Stems | 0–20 mm. |
0–20 mm. |
Leaves | 20–150(–400) × 10–120(–170) mm; petiole to 200 mm; blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute. |
100–300 × 80–200 mm; petiole to 300 mm; blade broadly oval to cordate-ovate, margins entire, veins conspicuous, laterals branching from midvein distal to base, surfaces glabrous. |
Scapes | 50–250(–500) mm, surpassing leaves or not, glabrous or hirsute. |
200–300 mm, glabrous. |
Spikes | brownish or greenish, (20–)50–300(–400) mm, densely flowered; bracts lanceolate, 0.5–1 mm, length 0.3–0.7 times sepals. |
brownish or greenish, 100–500 mm, loosely flowered, rachis visible between flowers; bracts round-ovate, 2 mm, length 0.8–1 times sepals. |
Flowers | sepals 1.5–2 mm; corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse; stamens 4. |
sepals 2–2.5 mm; corolla radially symmetric, lobes spreading, 2–2.5 mm, base obtuse; stamens 4. |
Fruits | (2–)4–5 mm, dehiscing at middle. |
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Seeds | 5–35, 0.5–1 mm. |
2–4, 2.5–3.5 mm. |
2n | = 12. |
= 24. |
Plantago major |
Plantago cordata |
|
Phenology | Flowering summer. | Flowering late spring–early summer. |
Habitat | Roadsides, trails, stream banks, urban areas, lawns, other disturbed areas. | Rocky or gravelly beds of shallow, slow-moving streams, sloughs, swamps. |
Elevation | 0–3000 m. (0–9800 ft.) | 0–200 m. (0–700 ft.) |
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; SPM; Greenland; Eurasia [Introduced in North America; introduced also in Mexico, Central America, South America, Africa, Pacific Islands, Australia]
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AL; AR; DC; FL; GA; IA; IL; IN; KY; MD; MI; MO; MS; NC; NY; OH; TN; VA; WI; ON
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Discussion | Subspecies intermedia (Gilibert) Lange is often accepted by European botanists as a separate species, Plantago uliginosa F. W. Schmidt. Observations suggest that plants referable to this taxon may occur in the United States; it has been reported from the New England states (A. Haines 2011). Subspecies intermedia is distinguished by more abundant (11–35) and smaller (0.8–1 mm) seeds, ascending spikes, ovoid fruits, and elliptic or lanceolate leaf blades. Without detailed morphologic and genetic investigations of North American plants similar to the study of European plants by M. Morgan-Richards and K. Wolff (1999), it is not possible to draw any conclusions about the status and distribution of this or any other possible infraspecific taxa of P. major in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plantago cordata is listed as federally endangered in Canada and is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 289. | FNA vol. 17, p. 285. |
Parent taxa | Plantaginaceae > Plantago | Plantaginaceae > Plantago |
Sibling taxa | ||
Synonyms | P. major var. pachyphylla, P. major var. pilgeri, P. major var. scopulorum | |
Name authority | Linnaeus: Sp. Pl. 1: 112. (1753) | Lamarck: in J. Lamarck and J. L. M. Poiret, Tabl. Encycl. 1: 338. (1792) |
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