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Alaska plantain, seashore plantain

blond plantain, desert Indian-wheat, desert plantain

Habit Perennials; caudex well developed, conspicuous, glabrous; roots fibrous, thick. Annuals; roots taproots, slender.
Stems

0–20 mm.

0–30 mm, often branched.

Leaves

(80–)100–400(–550) × (5–)10–35(–40) mm;

blade oblanceolate or almost linear, margins entire, veins conspicuous, 4 or 5, surfaces glabrous.

10–230 × 0.5–12 mm;

blade linear or narrowly elliptic, margins toothed, veins conspicuous or not, surfaces villous or lanate to sericeous.

Scapes

300–400 mm, glabrous or sparsely hairy, becoming densely so distally.

10–400 mm, hairy, hairs woolly, long.

Spikes

greenish or brownish, 350–450 mm, loosely flowered;

bracts ovate to deltate, 3–4 mm, length 1.5–2.5 times sepals.

grayish or brownish, 20–400 mm, densely flowered, flowers in spirals;

bracts ovate or elliptic, 1.7–4 mm, length 0.8–1.2 times sepals, apex not reached by green nerve.

Flowers

sepals 1.5–2 mm;

corolla radially symmetric, lobes spreading, 1.5–2 mm, base obtuse;

stamens 4.

sepals 1.9–3.5 mm;

corolla radially symmetric, lobes reflexed, 1.3–2.8 mm, base cuneate;

stamens 4.

Fruits

ovoid, indehiscent or dehiscence not circumscissile.

Seeds

1 or 2, 4–5 mm.

2, 2–2.6 mm.

2n

= 24.

= 8.

Plantago macrocarpa

Plantago ovata

Phenology Flowering late spring–early summer. Flowering spring.
Habitat Wet places, tidal marshes, saline areas. Sandy deserts and steppes.
Elevation 0–700 m. (0–2300 ft.) 0–1800 m. (0–5900 ft.)
Distribution
from FNA
AK; OR; WA; BC; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; TX; UT; Mexico (Baja California, Sonora); Eurasia; Africa
[WildflowerSearch map]
[BONAP county map]
Discussion

Plantago macrocarpa has been documented along the Pacific coast south to the mouth of the Yachats River in Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

From molecular evidence, S. C. Meyers and A. Liston (2008) suggested that Plantago ovata was introduced to North America during the Pleistocene. They recognized four varieties; North American specimens can be treated as two varieties based on bract and corolla color: the inland var. fastigiata (E. Morris) S. C. Meyers & Liston (midribs of mature flower bracts green, corolla lobes without reddish brown midribs) and the coastal var. insularis (Eastwood) S. C. Meyers & Liston (midribs of mature flower bracts brown, corolla lobe midribs prominent, reddish brown). Unfortunately, these features are not easily seen on many herbarium specimens, and these taxa are not recognized here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 289. FNA vol. 17, p. 290.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. insularis, P. insularis var. fastigiata
Name authority Chamisso & Schlechtendal: Linnaea 1: 166. (1826) Forsskål: Fl. Aegypt.-Arab., 31. (1775)
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