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Alaska plantain, seashore plantain

common plantain, great plantain, nippleseed, plantain majeur

Habit Perennials; caudex well developed, conspicuous, glabrous; roots fibrous, thick. Perennials, sometimes annuals; caudex absent; roots fibrous, thick.
Stems

0–20 mm.

0–20 mm.

Leaves

(80–)100–400(–550) × (5–)10–35(–40) mm;

blade oblanceolate or almost linear, margins entire, veins conspicuous, 4 or 5, surfaces glabrous.

20–150(–400) × 10–120(–170) mm;

petiole to 200 mm;

blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute.

Scapes

300–400 mm, glabrous or sparsely hairy, becoming densely so distally.

50–250(–500) mm, surpassing leaves or not, glabrous or hirsute.

Spikes

greenish or brownish, 350–450 mm, loosely flowered;

bracts ovate to deltate, 3–4 mm, length 1.5–2.5 times sepals.

brownish or greenish, (20–)50–300(–400) mm, densely flowered;

bracts lanceolate, 0.5–1 mm, length 0.3–0.7 times sepals.

Flowers

sepals 1.5–2 mm;

corolla radially symmetric, lobes spreading, 1.5–2 mm, base obtuse;

stamens 4.

sepals 1.5–2 mm;

corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse;

stamens 4.

Fruits

ovoid, indehiscent or dehiscence not circumscissile.

(2–)4–5 mm, dehiscing at middle.

Seeds

1 or 2, 4–5 mm.

5–35, 0.5–1 mm.

2n

= 24.

= 12.

Plantago macrocarpa

Plantago major

Phenology Flowering late spring–early summer. Flowering summer.
Habitat Wet places, tidal marshes, saline areas. Roadsides, trails, stream banks, urban areas, lawns, other disturbed areas.
Elevation 0–700 m. (0–2300 ft.) 0–3000 m. (0–9800 ft.)
Distribution
from FNA
AK; OR; WA; BC; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; SPM; Greenland; Eurasia [Introduced in North America; introduced also in Mexico, Central America, South America, Africa, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Plantago macrocarpa has been documented along the Pacific coast south to the mouth of the Yachats River in Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies intermedia (Gilibert) Lange is often accepted by European botanists as a separate species, Plantago uliginosa F. W. Schmidt. Observations suggest that plants referable to this taxon may occur in the United States; it has been reported from the New England states (A. Haines 2011). Subspecies intermedia is distinguished by more abundant (11–35) and smaller (0.8–1 mm) seeds, ascending spikes, ovoid fruits, and elliptic or lanceolate leaf blades. Without detailed morphologic and genetic investigations of North American plants similar to the study of European plants by M. Morgan-Richards and K. Wolff (1999), it is not possible to draw any conclusions about the status and distribution of this or any other possible infraspecific taxa of P. major in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 289. FNA vol. 17, p. 289.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. major var. pachyphylla, P. major var. pilgeri, P. major var. scopulorum
Name authority Chamisso & Schlechtendal: Linnaea 1: 166. (1826) Linnaeus: Sp. Pl. 1: 112. (1753)
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