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Alaska plantain, seashore plantain

Heller's plantain

Habit Perennials; caudex well developed, conspicuous, glabrous; roots fibrous, thick. Annuals; roots taproots, slender.
Stems

0–20 mm.

0–12 mm.

Leaves

(80–)100–400(–550) × (5–)10–35(–40) mm;

blade oblanceolate or almost linear, margins entire, veins conspicuous, 4 or 5, surfaces glabrous.

50–130 × 3–7 mm;

blade linear, margins entire, veins conspicuous or not, surfaces villous, adaxial rarely glabrous.

Scapes

300–400 mm, glabrous or sparsely hairy, becoming densely so distally.

80–250 mm, hairy, hairs patent, long and antrorse, short.

Spikes

greenish or brownish, 350–450 mm, loosely flowered;

bracts ovate to deltate, 3–4 mm, length 1.5–2.5 times sepals.

greenish, whitish, or brownish, 50–120 mm, densely flowered, flowers in whorls or pairs;

bracts triangular, 2.5–8 mm, length 0.6–2.2 times sepals.

Flowers

sepals 1.5–2 mm;

corolla radially symmetric, lobes spreading, 1.5–2 mm, base obtuse;

stamens 4.

sepals 3.5–4 mm;

corolla radially symmetric, lobes reflexed, 3–3.6 mm, base slightly cordate;

stamens 4, connective elongated, apex acute.

Fruits

ovoid, indehiscent or dehiscence not circumscissile.

Seeds

1 or 2, 4–5 mm.

2, 3.2–3.8 mm.

2n

= 24.

= 20.

Plantago macrocarpa

Plantago helleri

Phenology Flowering late spring–early summer. Flowering spring.
Habitat Wet places, tidal marshes, saline areas. Dry slopes and flats on limestone.
Elevation 0–700 m. (0–2300 ft.) 100–1700 m. (300–5600 ft.)
Distribution
from FNA
AK; OR; WA; BC; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
NM; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Plantago macrocarpa has been documented along the Pacific coast south to the mouth of the Yachats River in Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Populations of Plantago helleri are concentrated in central Texas, especially in the Edwards Plateau and Trans-Pecos. The species also occurs in Eddy and Lincoln counties, New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 289. FNA vol. 17, p. 287.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Name authority Chamisso & Schlechtendal: Linnaea 1: 166. (1826) Small: Bull. New York Bot. Gard. 1: 288. (1899)
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