Plantago indica |
Plantago rugelii |
|
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plantain des sables, sand plantain |
blackseed plantain, plantain de Rugel, Rugel's plantain |
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Habit | Annuals; roots taproots, slender. | Perennials, sometimes annuals; caudex absent; roots fibrous, thick. |
Stems | 100–350 mm, freely branched. |
0–20 mm. |
Leaves | cauline, opposite, 60–80 × 1–3 mm; blade linear to linear-lanceolate, margins entire or toothed, veins conspicuous or not, surfaces hairy. |
20–150 × 10–120 mm; petiole to 200 mm; blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute. |
Scapes | 100–650 mm, hairy. |
50–250 mm, glabrous or hirsute. |
Spikes | greenish or brownish, (50–)150–250 mm, densely flowered, eglandular; bracts proximal strongly differing from distal, ovate, 2–5(–7) mm, length 1–1.5 times sepals, proximal bracts: apex acute. |
brownish or greenish, 50–300 mm, densely or loosely flowered; bracts narrowly lanceolate, 2 mm, length 1–1.2 times sepals. |
Flowers | sepals 2–3 mm; corolla radially symmetric, lobes reflexed, 2–4 mm, base obtuse; stamens 4. |
sepals 1.5–2 mm; corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse; stamens 4. |
Fruits | 4–6(–8) mm, dehiscing proximal to middle. |
|
Seeds | 2, 2–2.5 mm. |
4 or 5(–8), 1.5–2 mm. |
2n | = 12. |
= 24. |
Plantago indica |
Plantago rugelii |
|
Phenology | Flowering late summer–fall. | Flowering summer. |
Habitat | Roadsides, railroads, sandy shorelines. | Open woods, meadows, pastures, waste places. |
Elevation | 0–200 m. (0–700 ft.) | 0–2000 m. (0–6600 ft.) |
Distribution |
CA; CT; DE; IA; IL; IN; MA; ME; MI; MN; MO; NC; NH; NJ; NY; OH; OR; PA; VA; VT; WA; WI; BC; MB; ON; QC; Eurasia [Introduced in North America] |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
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Discussion | While considering the proposal made by W. L. Applequist (2006) to reject Plantago psyllium, the Nomenclature Committee for Vascular Plants also decided that P. indica is a legitimate name (R. K. Brummitt 2009). A. B. Doweld and A. Shipunov (2017) published a proposal to reject P. indica in favor of P. arenaria. That proposal awaits a decision by that committee. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants of Plantago major and P. rugelii are morphologically indistinguishable when young; they may be differentiated with certainty only at fruiting stage. DNA sequence data (A. Shipunov et al. 2014) confirm that the two species are distinct: P. rugelii is different from P. major by 11 substitutions in the ITS2 sequence. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 288. | FNA vol. 17, p. 292. |
Parent taxa | Plantaginaceae > Plantago | Plantaginaceae > Plantago |
Sibling taxa | ||
Synonyms | P. arenaria, P. psyllium | |
Name authority | Linnaeus: Syst. Nat., ed. 10, 2: 896. 1759, legitimacy of — name in question | Decaisne: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(1): 700. (1852) |
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